Hawaiian Insects
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Banza nitida TPR 9684
Banza nitida, a native flightless katydid. Tree Planting Road.

Banza nitida TPR 9685
Banza nitida, a native flightless katydid. Tree Planting Road.

Banza nitida TPR 9687
Banza nitida, a native flightless katydid. Tree Planting Road.

koa defoliation mauka 3576
The southern extent of koa defoliation seen in the distance from the Saddle...
The southern extent of koa defoliation seen in the distance from the Saddle Road, near the 15 mile marker. There seem to be relatively few places where it can be seen from the ground, due to the topography and distance from roads and trails.

Lispocephala Saddle 2447
October 4, 2009. Tetraplasandra kipuka, north of Kaumana trailhead....
October 4, 2009. Tetraplasandra kipuka, north of Kaumana trailhead. Lispocephala sp.

Lispocephala Saddle 2451
October 4, 2009. Tetraplasandra kipuka, north of Kaumana trailhead....
October 4, 2009. Tetraplasandra kipuka, north of Kaumana trailhead. Lispocephala sp.

Lispocephala Saddle 2452
October 4, 2009. Tetraplasandra kipuka, north of Kaumana trailhead....
October 4, 2009. Tetraplasandra kipuka, north of Kaumana trailhead. Lispocephala sp.

Drosophila tanythrix Kipuka 14 2606
October 6, 2009. Kipuka 14, south of of Kaumana Trail. Drosophila tanythrix...
October 6, 2009. Kipuka 14, south of of Kaumana Trail. Drosophila tanythrix mating.

Drosophila tanythrix Kipuka 14 2605
October 6, 2009. Kipuka 14, south of of Kaumana Trail. Drosophila tanythrix...
October 6, 2009. Kipuka 14, south of of Kaumana Trail. Drosophila tanythrix mating.

Drosophila tanythrix Kipuka 14 2604
October 6, 2009. Kipuka 14, south of of Kaumana Trail. Drosophila tanythrix...
October 6, 2009. Kipuka 14, south of of Kaumana Trail. Drosophila tanythrix mating.

Drosophila mulli Saddle 0946
Pupal case of Drosophila mulli under leaves of Pritchardia beccariana palm,...
Pupal case of Drosophila mulli under leaves of Pritchardia beccariana palm, loulu. Off of Saddle Road. This would seem to show fairly clearly that they're breeding in the Pritchardia, though it's not clear exactly where. Perhaps in the rachis when it first dies and rots but before it falls down (by which time it's usually too decayed for typical Drosophila tastes), from where the larvae crawl out onto the lowest living leaf to pupate. No pupae were seen on upper leaves.

Drosophila mulli Saddle 0947
Pupal case of Drosophila mulli under leaves of Pritchardia beccariana palm,...
Pupal case of Drosophila mulli under leaves of Pritchardia beccariana palm, loulu. Off of Saddle Road. This would seem to show fairly clearly that they're breeding in the Pritchardia, though it's not clear exactly where. Perhaps in the rachis when it first dies and rots but before it falls down (by which time it's usually too decayed for typical Drosophila tastes), from where the larvae crawl out onto the lowest living leaf to pupate. No pupae were seen on upper leaves.

Drosophila mulli Saddle 0948
Drosophila mulli under leaves of Pritchardia beccariana palm, loulu. Off of...
Drosophila mulli under leaves of Pritchardia beccariana palm, loulu. Off of Saddle Road.

Drosophila mulli under leaves of Pritchardia beccariana palm, loulu. Off of...
Drosophila mulli under leaves of Pritchardia beccariana palm, loulu. Off of Saddle Road.

Asteia hawaiiensis Saddle 0968
Asteia hawaiiensis under leaf of Pritchardia beccariana palm, loulu. Off of...
Asteia hawaiiensis under leaf of Pritchardia beccariana palm, loulu. Off of Saddle Road.

Asteia nr sabroskyi Saddle 0962
Asteia nr. sabroskyi on a rachis of Pritchardia beccariana palm, loulu. Off...
Asteia nr. sabroskyi on a rachis of Pritchardia beccariana palm, loulu. Off of Saddle Road.

Hyposmocoma sp Saddle 0954
Case of a Hyposmocoma caterpillar under a leaf of Pritchardia beccariana...
Case of a Hyposmocoma caterpillar under a leaf of Pritchardia beccariana palm, loulu. Off of Saddle Road.

Udara blackburni Saddle 0937
Caterpillar of Udara blackburni, Blackburn's blue butterfly, feeding on koa...
Caterpillar of Udara blackburni, Blackburn's blue butterfly, feeding on koa leaves. Off of Saddle Road.

Udara blackburni Saddle 0938
Caterpillar of Udara blackburni, Blackburn's blue butterfly, feeding on koa...
Caterpillar of Udara blackburni, Blackburn's blue butterfly, feeding on koa leaves. Off of Saddle Road.

Nesodynerus rubopustulatus Saddle 9139
Nesodynerus, nalo hope eha. Saddle Road, 1855 lava flow, above 20 mile mark.

Nesodynerus scoriaceus Saddle 9130
Nesodynerus scoriaceus, nalo hope eha. Saddle Road, 1855 lava flow, above 20...
Nesodynerus scoriaceus, nalo hope eha. Saddle Road, 1855 lava flow, above 20 mile mark.

Nesodynerus scoriaceus Saddle 9131
Nesodynerus scoriaceus, nalo hope eha. Saddle Road, 1855 lava flow, above 20...
Nesodynerus scoriaceus, nalo hope eha. Saddle Road, 1855 lava flow, above 20 mile mark.

Nesodynerus scoriaceus Saddle 9134
Nesodynerus scoriaceus, nalo hope eha. Saddle Road, 1855 lava flow, above 20...
Nesodynerus scoriaceus, nalo hope eha. Saddle Road, 1855 lava flow, above 20 mile mark.

Nesodynerus scoriaceus Saddle 9140a

Nesodynerus scoriaceus Saddle 9141a

Nesodynerus scoriaceus Saddle 9143a

Nesodynerus spp Saddle 9138
Nesodynerus rubopustulatus (left) and N. scoriaceus (right), nalo hope eha....
Nesodynerus rubopustulatus (left) and N. scoriaceus (right), nalo hope eha. Saddle Road, 1855 lava flow, above 20 mile mark.

Theridion grallator Kulani 8111
Kulani, Puu Makaala NAR. Theridion grallator, happyface spider female with...
Kulani, Puu Makaala NAR. Theridion grallator, happyface spider female with egg sac.

Drosophila basisetae Stainback 0430
July 21, 2009. Stainback Highway, 3600' diagonal road. Drosophila basisetae.

Drosophila percnosoma Stainback 0380
July 21, 2009. Stainback Highway, 3600' diagonal road. Drosophila percnosoma.

Drosophila prolaticilia Stainback 0460
July 21, 2009. Stainback Highway, 3600' diagonal road. Drosophila...
July 21, 2009. Stainback Highway, 3600' diagonal road. Drosophila prolaticilia.

Drosophila prolaticilia Stainback 0461
July 21, 2009. Stainback Highway, 3600' diagonal road. Drosophila...
July 21, 2009. Stainback Highway, 3600' diagonal road. Drosophila prolaticilia.

Drosophila prolaticilia Stainback 0462
July 21, 2009. Stainback Highway, 3600' diagonal road. Drosophila...
July 21, 2009. Stainback Highway, 3600' diagonal road. Drosophila prolaticilia.

Drosophila setosimentum Stainback 0421
July 21, 2009. Stainback Highway, 3600' diagonal road. Drosophila setosimentum.

Drosophila setosimentum Stainback 0422
July 21, 2009. Stainback Highway, 3600' diagonal road. Drosophila setosimentum.

Drosophila setosimentum Stainback 0426
July 21, 2009. Stainback Highway, 3600' diagonal road. Drosophila setosimentum.

Drosophila spp Stainback 0429
July 21, 2009. Stainback Highway, 3600' diagonal road. L-R: D. sproati and...
July 21, 2009. Stainback Highway, 3600' diagonal road. L-R: D. sproati and D. basisetae.

Drosophila spp Stainback 0431
July 21, 2009. Stainback Highway, 3600' diagonal road. L-R: Drosophila...
July 21, 2009. Stainback Highway, 3600' diagonal road. L-R: Drosophila prolaticilia, D. sproati, and D. basisetae.

Drosophila spp Stainback 0433
July 21, 2009. Stainback Highway, 3600' diagonal road. L-R: Drosophila...
July 21, 2009. Stainback Highway, 3600' diagonal road. L-R: Drosophila prolaticilia, D. sproati, and D. basisetae.

Drosophila sproati Stainback 0382
July 21, 2009. Stainback Highway, 3600' diagonal road. Drosophila sproati.

Drosophila sproati Stainback 0385
July 21, 2009. Stainback Highway, 3600' diagonal road. Drosophila sproati.

Drosophila sproati Stainback 0393
July 21, 2009. Stainback Highway, 3600' diagonal road. Drosophila sproati.

Scaptomyza cyrtandrae Stainback 0448
July 21, 2009. Stainback Highway, 3600' diagonal road. Scaptomyza...
July 21, 2009. Stainback Highway, 3600' diagonal road. Scaptomyza cyrtandrae on Cyrtandra platyphylla.

Scaptomyza cyrtandrae Stainback 0449
July 21, 2009. Stainback Highway, 3600' diagonal road. Scaptomyza...
July 21, 2009. Stainback Highway, 3600' diagonal road. Scaptomyza cyrtandrae on Cyrtandra platyphylla.

Scaptomyza cyrtandrae Stainback 0451
July 21, 2009. Stainback Highway, 3600' diagonal road. Scaptomyza...
July 21, 2009. Stainback Highway, 3600' diagonal road. Scaptomyza cyrtandrae on Cyrtandra platyphylla.

Scaptomyza cyrtandrae

Drosophila prolaticilia R Road1
June 21, 2009. R Road, Stainback Highway. Drosophila prolaticilia feeding...
June 21, 2009. R Road, Stainback Highway. Drosophila prolaticilia feeding on rotting Clermontia parviflora branch. This was the most common picture wing species in the area, despite there being little of its only known host, Freycinetia; only one D. sproati was seen.

Drosophila prolaticilia R Road2
June 21, 2009. R Road, Stainback Highway. Drosophila prolaticilia feeding...
June 21, 2009. R Road, Stainback Highway. Drosophila prolaticilia feeding on rotting Clermontia parviflora branch. This was the most common picture wing species in the area, despite there being little of its only known host, Freycinetia; only one D. sproati was seen.

Drosophila prolaticilia R Road3
June 21, 2009. R Road, Stainback Highway. Drosophila prolaticilia feeding...
June 21, 2009. R Road, Stainback Highway. Drosophila prolaticilia feeding on rotting Clermontia parviflora branch. This was the most common picture wing species in the area, despite there being little of its only known host, Freycinetia; only one D. sproati was seen.

Drosophila ochracea Stainback 3617
January 12, 2009. Stainback Highway, 3600'. Drosophila ochracea.

Drosophila ochracea Stainback 3618
January 12, 2009. Stainback Highway, 3600'. Drosophila ochracea.

Drosophila ochracea Stainback 3621
January 12, 2009. Stainback Highway, 3600'. Drosophila ochracea.

January 12, 2009. Stainback Highway, 3600'. Drosophila setosimentum.

Tetragnatha Stainback 3636
January 12, 2009. Stainback Highway, 3600'. Tetragnatha sp.

Orsonwelles graphicus R Road 2728
October 24, 2009. Tom's Trail (R Road), Stainback Highway. Orsonwelles...
October 24, 2009. Tom's Trail (R Road), Stainback Highway. Orsonwelles graphicus. It's very rare to see these out during the day, though the webs are abundant throughout the forest.

Orsonwelles graphicus R Road 2729
October 24, 2009. Tom's Trail (R Road), Stainback Highway. Orsonwelles...
October 24, 2009. Tom's Trail (R Road), Stainback Highway. Orsonwelles graphicus. It's very rare to see these out during the day, though the webs are abundant throughout the forest.

Orsonwelles graphicus R Road 2733
October 24, 2009. Tom's Trail (R Road), Stainback Highway. Orsonwelles...
October 24, 2009. Tom's Trail (R Road), Stainback Highway. Orsonwelles graphicus. It's very rare to see these out during the day, though the webs are abundant throughout the forest.

November 22, 2009. Tom's Trail (R Road), Stainback Highway. Orsonwelles...
November 22, 2009. Tom's Trail (R Road), Stainback Highway. Orsonwelles graphicus male.

Drosophila prolaticilia R Road 2490
October 5, 2009. Tom's Trail (R Road), Stainback Highway. Drosophila...
October 5, 2009. Tom's Trail (R Road), Stainback Highway. Drosophila prolaticilia.

Tetragnatha R Road 2507
October 5, 2009. Tom's Trail (R Road), Stainback Highway. Tetragnatha sp.,...
October 5, 2009. Tom's Trail (R Road), Stainback Highway. Tetragnatha sp., feeding on Drosophila percnosoma.

Tetragnatha R Road 2509
October 5, 2009. Tom's Trail (R Road), Stainback Highway. Tetragnatha sp.,...
October 5, 2009. Tom's Trail (R Road), Stainback Highway. Tetragnatha sp., feeding on Drosophila percnosoma.

Tetragnatha R Road 2510
October 5, 2009. Tom's Trail (R Road), Stainback Highway. Tetragnatha sp.,...
October 5, 2009. Tom's Trail (R Road), Stainback Highway. Tetragnatha sp., feeding on Drosophila percnosoma.

Drosophila ochracea R Road 2477
October 5, 2009. Tom's Trail (R Road), Stainback Highway. Drosophila ochracea.

Drosophila ochracea R Road 2478
October 5, 2009. Tom's Trail (R Road), Stainback Highway. Drosophila ochracea.

Drosophila ochracea R Road 2479
October 5, 2009. Tom's Trail (R Road), Stainback Highway. Drosophila ochracea.

Drosophila pullipes Army Road 0693
Puu Makaala NAR, Army Road Makai. Drosophila pullipes on a rotting branch of...
Puu Makaala NAR, Army Road Makai. Drosophila pullipes on a rotting branch of Wikstroemia sandwicensis, its host.

Drosophila pullipes Army Road 0694
Puu Makaala NAR, Army Road Makai. Drosophila pullipes on a rotting branch of...
Puu Makaala NAR, Army Road Makai. Drosophila pullipes on a rotting branch of Wikstroemia sandwicensis, its host.

Drosophila pullipes Army Road 0695
Puu Makaala NAR, Army Road Makai. Drosophila pullipes on a rotting branch of...
Puu Makaala NAR, Army Road Makai. Drosophila pullipes on a rotting branch of Wikstroemia sandwicensis, its host.

Drosophila pullipes Army Road 6233 crop
Puu Makaala NAR, Army Road Makai. Drosophila pullipes on fallen Wikstroemia...
Puu Makaala NAR, Army Road Makai. Drosophila pullipes on fallen Wikstroemia sandwicensis tree.

Puu Makaala NAR, Army Road Makai. Drosophila pullipes on fallen Wikstroemia...
Puu Makaala NAR, Army Road Makai. Drosophila pullipes on fallen Wikstroemia sandwicensis tree.

Puu Makaala NAR, Army Road Makai. Drosophila pullipes on fallen Wikstroemia sandwicensis tree. The fly took off as I took the picture, doing a back flip.

December 8, 2009. Stainback Highway, across from Army Road Makai....
December 8, 2009. Stainback Highway, across from Army Road Makai. Drosophila setosimentum.

December 8, 2009. Stainback Highway, across from Army Road Makai....
December 8, 2009. Stainback Highway, across from Army Road Makai. Drosophila sproati.

December 8, 2009. Stainback Highway, across from Army Road Makai....
December 8, 2009. Stainback Highway, across from Army Road Makai. Scaptomyza cyrtandrae on Cyrtandra platyphylla.

December 8, 2009. Stainback Highway, across from Army Road Makai. Scaptomyza cyrtandrae on Cyrtandra platyphylla.

Elimaea punctifera Olaa Flume 9213
Elimaea punctifera, an alien katydid. Olaa Flume Road. This is a nymph;...
Elimaea punctifera, an alien katydid. Olaa Flume Road. This is a nymph; they are distinctive for their habit of standing with the head down and hind legs splayed apart.

Elimaea punctifera Olaa Flume 9214
Elimaea punctifera, an alien katydid. Olaa Flume Road. This is a nymph;...
Elimaea punctifera, an alien katydid. Olaa Flume Road. This is a nymph; they are distinctive for their habit of standing with the head down and hind legs splayed apart.

Elimaea punctifera Olaa Flume 9217
Elimaea punctifera, an alien katydid. Olaa Flume Road. This is a nymph;...
Elimaea punctifera, an alien katydid. Olaa Flume Road. This is a nymph; they are distinctive for their habit of standing with the head down and hind legs splayed apart.

Nesodynerus scoriaceus. Kipuka Ainahou, Saddle Road.

Nesodynerus sp Saddle 9192
Nesodynerus sp. Kipuka Ainahou, Saddle Road.

Sierola sp Saddle 9197
Sierola sp. Kipuka Ainahou, Saddle Road.

Sierola sp Saddle 9198
Sierola sp. Kipuka Ainahou, Saddle Road.

Hylaeus difficilis Puu Kole 5573
July 10, 2010. Mauna Kea, near Puu Kole. Hylaeus difficilis visiting...
July 10, 2010. Mauna Kea, near Puu Kole. Hylaeus difficilis visiting flowers of Geranium cuneatum.

Hylaeus difficilis Puu Kole 5574
July 10, 2010. Mauna Kea, near Puu Kole. Hylaeus difficilis visiting...
July 10, 2010. Mauna Kea, near Puu Kole. Hylaeus difficilis visiting flowers of Geranium cuneatum.

Hylaeus difficilis Puu Kole 5577
July 10, 2010. Mauna Kea, near Puu Kole. Hylaeus difficilis visiting...
July 10, 2010. Mauna Kea, near Puu Kole. Hylaeus difficilis visiting flowers of Geranium cuneatum.

Hylaeus difficilis Hakalau 7519
Hakalau NWR. Hylaeus difficilis at a nesting site.

Hylaeus difficilis Hakalau 7520
Hakalau NWR. Hylaeus difficilis at a nesting site.

Hylaeus sphecodoides Hakalau 7528
Hakalau NWR. Hylaeus sphecodoides, a cleptoparasitic species.

Hylaeus sphecodoides Hakalau 7530
Hakalau NWR. Hylaeus sphecodoides, a cleptoparasitic species.

Hylaeus sphecodoides Hakalau 7531
Hakalau NWR. Hylaeus sphecodoides, a cleptoparasitic species.

Hylaeus difficilis Hakalau 7557
Hakalau NWR. Hylaeus difficilis visiting a naio flower.

Hakalau NWR. Hylaeus difficilis visiting a naio flower.

Udara blackburni Hakalau 7539
Hakalau NWR. Udara blackburni, one of the two native butterflies, visiting a...
Hakalau NWR. Udara blackburni, one of the two native butterflies, visiting a naio flower.

Udara blackburni Hakalau 7540
Hakalau NWR. Udara blackburni, one of the two native butterflies, visiting a...
Hakalau NWR. Udara blackburni, one of the two native butterflies, visiting a naio flower.

Udara blackburni Hakalau 7541
Hakalau NWR. Udara blackburni, one of the two native butterflies, visiting a...
Hakalau NWR. Udara blackburni, one of the two native butterflies, visiting a naio flower.

Trupanea arboreae Waipahoehoe 5692
July 11, 2010. Wailuku plant sanctuary, Waipahoehoe Gulch. Trupanea...
July 11, 2010. Wailuku plant sanctuary, Waipahoehoe Gulch. Trupanea arboreae on Argyroxiphium sandwicense.

Trupanea arboreae Waipahoehoe 5702
July 11, 2010. Wailuku plant sanctuary, Waipahoehoe Gulch. Trupanea...
July 11, 2010. Wailuku plant sanctuary, Waipahoehoe Gulch. Trupanea arboreae on Argyroxiphium sandwicense.

Hylaeus difficilis Waipahoehoe 5667
July 11, 2010. Wailuku plant sanctuary, Waipahoehoe Gulch. Hylaeus...
July 11, 2010. Wailuku plant sanctuary, Waipahoehoe Gulch. Hylaeus difficilis on apparent Dubautia arborea x linearis hybrid.

Hylaeus difficilis Waipahoehoe 5673
July 11, 2010. Wailuku plant sanctuary, Waipahoehoe Gulch. Hylaeus...
July 11, 2010. Wailuku plant sanctuary, Waipahoehoe Gulch. Hylaeus difficilis on apparent Dubautia arborea x linearis hybrid.

Hylaeus difficilis Waipahoehoe 5674
July 11, 2010. Wailuku plant sanctuary, Waipahoehoe Gulch. Hylaeus...
July 11, 2010. Wailuku plant sanctuary, Waipahoehoe Gulch. Hylaeus difficilis on apparent Dubautia arborea x linearis hybrid.

July 11, 2010. Wailuku plant sanctuary, Waipahoehoe Gulch. Hylaeus difficilis on Dubautia ciliolata subsp. glutinosa.

Trupanea arboreae Hale Pohaku 5555
July 10, 2010. Hale Pohaku. Trupanea arboreae pupae in seed heads of Mauna...
July 10, 2010. Hale Pohaku. Trupanea arboreae pupae in seed heads of Mauna Kea silversword, Argyroxiphium sandwicense subsp. sandwicense. Seed heads here were heavily infested, with five or more old pupae in each, despite there being only a few plants. At the larger Waipahoehoe population there was much less damage.

Trupanea arboreae Hale Pohaku 5558
July 10, 2010. Hale Pohaku. Trupanea arboreae pupae in seed heads of Mauna...
July 10, 2010. Hale Pohaku. Trupanea arboreae pupae in seed heads of Mauna Kea silversword, Argyroxiphium sandwicense subsp. sandwicense. Seed heads here were heavily infested, with five or more old pupae in each, despite there being only a few plants. At the larger Waipahoehoe population there was much less damage.

Trupanea arboreae Hale Pohaku 5563
July 10, 2010. Hale Pohaku. Trupanea arboreae on Mauna Kea silversword,...
July 10, 2010. Hale Pohaku. Trupanea arboreae on Mauna Kea silversword, Argyroxiphium sandwicense subsp. sandwicense. These are outplanted near the visitor center.

July 10, 2010. Hale Pohaku. Trupanea arboreae on Mauna Kea silversword,...
July 10, 2010. Hale Pohaku. Trupanea arboreae on Mauna Kea silversword, Argyroxiphium sandwicense subsp. sandwicense. These are outplanted near the visitor center.

Lispocephala philydra Kolekole 9317
Lispocephala philydra, an endemic predatory fly. Kolekole stream.

Sigmatineurum omega Kolekole 9315
Sigmatineurum omega, Kolekole stream.

Drosophila digressa Olaa 3510
January 1, 2010. Hawaii Volcanoes NP, Olaa Small Tract, Pole 44. Drosophila...
January 1, 2010. Hawaii Volcanoes NP, Olaa Small Tract, Pole 44. Drosophila digressa.

Drosophila digressa Olaa 3511
January 1, 2010. Hawaii Volcanoes NP, Olaa Small Tract, Pole 44. Drosophila...
January 1, 2010. Hawaii Volcanoes NP, Olaa Small Tract, Pole 44. Drosophila digressa.

Drosophila digressa Olaa 3516
January 1, 2010. Hawaii Volcanoes NP, Olaa Small Tract, Pole 44. Drosophila...
January 1, 2010. Hawaii Volcanoes NP, Olaa Small Tract, Pole 44. Drosophila digressa.

Drosophila macrothrix Olaa 3541
January 1, 2010. Hawaii Volcanoes NP, Olaa Small Tract, Pole 44. Drosophila macrothrix, with D. sproati on the left.

Drosophila macrothrix Olaa 3544
January 1, 2010. Hawaii Volcanoes NP, Olaa Small Tract, Pole 44. Drosophila macrothrix.

Drosophila macrothrix Olaa 7140
Hawaii Volcanoes NP, Olaa Small Tract, Pole 44. Drosophila macrothrix.

Hawaii Volcanoes NP, Olaa Small Tract, Pole 44. Drosophila macrothrix.

Drosophila paucipuncta Olaa 3521
January 1, 2010. Hawaii Volcanoes NP, Olaa Small Tract, Pole 44. Drosophila paucipuncta.

Drosophila paucipuncta Olaa 3524
January 1, 2010. Hawaii Volcanoes NP, Olaa Small Tract, Pole 44. Drosophila paucipuncta.

Drosophila paucipuncta Olaa 3528
January 1, 2010. Hawaii Volcanoes NP, Olaa Small Tract, Pole 44. Drosophila paucipuncta.

Drosophila percnosoma Olaa 7127
Hawaii Volcanoes NP, Olaa Small Tract, Pole 44. Drosophila percnosoma males...
Hawaii Volcanoes NP, Olaa Small Tract, Pole 44. Drosophila percnosoma males competing for lek positions.

Drosophila yooni Olaa 7130
Hawaii Volcanoes NP, Olaa Small Tract, Pole 44. Drosophila yooni.

Drosophila yooni Olaa 7133
Hawaii Volcanoes NP, Olaa Small Tract, Pole 44. Drosophila yooni.

Lispocephala sp Olaa 3545
January 1, 2010. Hawaii Volcanoes NP, Olaa Small Tract, Pole 44....
January 1, 2010. Hawaii Volcanoes NP, Olaa Small Tract, Pole 44. Lispocephala sp.

Lispocephala sp Olaa 3546
January 1, 2010. Hawaii Volcanoes NP, Olaa Small Tract, Pole 44....
January 1, 2010. Hawaii Volcanoes NP, Olaa Small Tract, Pole 44. Lispocephala sp.

Lispocephala sp Olaa 3548
January 1, 2010. Hawaii Volcanoes NP, Olaa Small Tract, Pole 44....
January 1, 2010. Hawaii Volcanoes NP, Olaa Small Tract, Pole 44. Lispocephala sp. The movement is a result of it reacting to the sound of the mirror flipping up before the shutter opened, showing how fast its reflexes are.

January 1, 2010. Hawaii Volcanoes NP, Olaa Small Tract, Pole 44. Lispocephala sp. The movement is a result of it reacting to the sound of the mirror flipping up before the shutter opened, showing how fast its reflexes are.

Lycosa howarthi Puaulu cave 1274
Lycosa howarthi, the "little-eyed big-eyed spider". Lava tube cave, Kipuka...
Lycosa howarthi, the "little-eyed big-eyed spider". Lava tube cave, Kipuka Puaulu.

Lycosa howarthi Puaulu cave 1275
Lycosa howarthi, the "little-eyed big-eyed spider". Lava tube cave, Kipuka...
Lycosa howarthi, the "little-eyed big-eyed spider". Lava tube cave, Kipuka Puaulu.

Lycosa howarthi Puaulu cave 1280
Lycosa howarthi, the "little-eyed big-eyed spider". Lava tube cave, Kipuka...
Lycosa howarthi, the "little-eyed big-eyed spider". Lava tube cave, Kipuka Puaulu.

Mashuri Puaulu cave
Mashuri checking out the coating on the wall of the lava tube cave, Kipuka...
Mashuri checking out the coating on the wall of the lava tube cave, Kipuka Puaulu.

Oliarus polyphemus Puaulu cave 1286
Oliarus polyphemus, a planthopper that inhabits caves, feeding on ohia roots...
Oliarus polyphemus, a planthopper that inhabits caves, feeding on ohia roots that penetrate the ceiling. Lava tube cave, Kipuka Puaulu.

Thaumatogryllus cavicola Puaulu cave 1284
Thaumatogryllus cavicola, an endemic cave cricket. Lava tube cave, Kipuka...
Thaumatogryllus cavicola, an endemic cave cricket. Lava tube cave, Kipuka Puaulu.

Thaumatogryllus cavicola Puaulu cave 1285
Thaumatogryllus cavicola, an endemic cave cricket. Lava tube cave, Kipuka...
Thaumatogryllus cavicola, an endemic cave cricket. Lava tube cave, Kipuka Puaulu.

tipulid tracks Puaulu cave 1289
Tracks made in the microbial coating of the rock walls, apparently by crane...
Tracks made in the microbial coating of the rock walls, apparently by crane fly larvae. Lava tube cave, Kipuka Puaulu.

tipulid tracks Puaulu cave 1292
Tracks made in the microbial coating of the rock walls, apparently by crane...
Tracks made in the microbial coating of the rock walls, apparently by crane fly larvae. Lava tube cave, Kipuka Puaulu.

tipulid tracks Puaulu cave 1297
Tracks made in the microbial coating of the rock walls, apparently by crane...
Tracks made in the microbial coating of the rock walls, apparently by crane fly larvae. Lava tube cave, Kipuka Puaulu.

Hyposmocoma parasitoid Olaa 4977
May 2, 2010. Olaa Forest Reserve, behind Volcano Transfer Station....
May 2, 2010. Olaa Forest Reserve, behind Volcano Transfer Station. Chalcidioid parasite attacking a Hyposmocoma caterpillar.

Olaa Forest Reserve, behind Volcano transfer station. Drosophila paucipuncta.

Drosophila prolaticilia Olaa 6112
Olaa Forest Reserve, behind Volcano transfer station. Drosophila prolaticilia.

Drosophila prolaticilia Olaa 6113
Olaa Forest Reserve, behind Volcano transfer station. Drosophila prolaticilia.

Drosophila prolaticilia Olaa 6116
Olaa Forest Reserve, behind Volcano transfer station. Drosophila prolaticilia.

Drosophila tetraplasandrae Olaa 6121
Olaa Forest Reserve, behind Volcano transfer station. Drosophila...
Olaa Forest Reserve, behind Volcano transfer station. Drosophila tetraplasandrae.

Drosophila tetraplasandrae Olaa 6131
Olaa Forest Reserve, behind Volcano transfer station. Drosophila...
Olaa Forest Reserve, behind Volcano transfer station. Drosophila tetraplasandrae.

Drosophila tetraplasandrae Olaa 6132
Olaa Forest Reserve, behind Volcano transfer station. Drosophila...
Olaa Forest Reserve, behind Volcano transfer station. Drosophila tetraplasandrae.

house spider Puaulu 3565
January 1, 2010. Hawaii Volcanoes NP, Kipuka Puaulu. Webs of the Australian...
January 1, 2010. Hawaii Volcanoes NP, Kipuka Puaulu. Webs of the Australian house spider (Badumna longinqua?), which David Foote thinks is responsible for wiping out Drosophila engyochracea in Kipuka Ki and Puaulu starting in 2005. It is particularly fond of ohia trees with their large pieces of peeling bark.

house spider Puaulu 3564
January 1, 2010. Hawaii Volcanoes NP, Kipuka Puaulu. Webs of the Australian...
January 1, 2010. Hawaii Volcanoes NP, Kipuka Puaulu. Webs of the Australian house spider (Badumna longinqua?), which David Foote thinks is responsible for wiping out Drosophila engyochracea in Kipuka Ki and Puaulu starting in 2005. It is particularly fond of ohia trees with their large pieces of peeling bark.

Lycosa sp Mauna Kea 8593
Lycosa sp., near the summit of Mauna Kea.

Nysius wekiuicola Mauna Kea 8600
Nysius wekiuicola, the wekiu bug. Puu Wekiu, Mauna Kea.

Nysius wekiuicola Mauna Kea 8603
Nysius wekiuicola, the wekiu bug. Puu Wekiu, Mauna Kea.

Nysius wekiuicola Mauna Kea 8606
Nysius wekiuicola, the wekiu bug. Puu Wekiu, Mauna Kea.

Telmatogeton torrenticola Kolekole 3472
Habitat of Telmatogeton torrenticola, a very large endemic midge. Umauma...
Habitat of Telmatogeton torrenticola, a very large endemic midge. Umauma stream. These flies live in fast-flowing streams, rarely flying and unperturbed by falling into the water. Large numbers were visible running over the rocks in mating swarms, and the larvae could also be seen crawling around feeding on algae.

Telmatogeton torrenticola Kolekole 9301
Telmatogeton torrenticola, a very large endemic midge. Umauma stream. These...
Telmatogeton torrenticola, a very large endemic midge. Umauma stream. These flies live in fast-flowing streams, rarely flying and unperturbed by falling into the water. Large numbers were visible running over the rocks in mating swarms, and the larvae could also be seen crawling around feeding on algae.

Telmatogeton torrenticola Kolekole 9303
Telmatogeton torrenticola, a very large endemic midge. Umauma stream. These...
Telmatogeton torrenticola, a very large endemic midge. Umauma stream. These flies live in fast-flowing streams, rarely flying and unperturbed by falling into the water. Large numbers were visible running over the rocks in mating swarms, and the larvae could also be seen crawling around feeding on algae.

Telmatogeton torrenticola, a very large endemic midge. Umauma stream. These...
Telmatogeton torrenticola, a very large endemic midge. Umauma stream. These flies live in fast-flowing streams, rarely flying and unperturbed by falling into the water. Large numbers were visible running over the rocks in mating swarms, and the larvae could also be seen crawling around feeding on algae.

Nysius wekiuicola Mauna Kea death trap 8581
A shaft used for holding flags in the snow in a parking lot at the telescopes...
A shaft used for holding flags in the snow in a parking lot at the telescopes on Mauna Kea, and which serves as a permanent pitfall trap for wekiu bugs.

Nysius wekiuicola Mauna Kea death trap 8583
Looking inside the tube in the Mauna Kea telescope parking lot, with hundreds...
Looking inside the tube in the Mauna Kea telescope parking lot, with hundreds of dead wekiu bugs in the bottom. The rotting ones in there probably continue to attract others.

A pile of wekiu bugs (Nysius wekiuicola) recently freed from a live trap near...
A pile of wekiu bugs (Nysius wekiuicola) recently freed from a live trap near the Keck telescopes.

koa defoliation Waikaumalo 9347
A patch of koa partially defoliated by Scotorythra paludicola, koa moth....
A patch of koa partially defoliated by Scotorythra paludicola, koa moth. Visible from about 1900 ft elevation, Waikaumalo stream, just below the main koa defoliation area.

Scotorythra paludicola feeding Waikaumalo 9333
Leaf damage (epidermis scraping) by Scotorythra paludicola, koa moth....
Leaf damage (epidermis scraping) by Scotorythra paludicola, koa moth. Waikaumalo stream, below the koa defoliation area.

Scotorythra paludicola feeding Waikaumalo 9334
Leaf damage (epidermis scraping) by Scotorythra paludicola, koa moth....
Leaf damage (epidermis scraping) by Scotorythra paludicola, koa moth. Waikaumalo stream, below the koa defoliation area.

Drosophila hawaiiensis Laupahoehoe 7182
Laupahoehoe NAR. Drosophila hawaiiensis.

Drosophila hawaiiensis Laupahoehoe 7185
Laupahoehoe NAR. Drosophila hawaiiensis.

Drosophila hawaiiensis Laupahoehoe 7195
Laupahoehoe NAR. Drosophila hawaiiensis.

Theridion grallator black Thurston 1740
Escape Rd. near Thurston lava tube, Hawaii Volcanoes National Park....
Escape Rd. near Thurston lava tube, Hawaii Volcanoes National Park. Theridion grallator, happyface spider.

Theridion grallator black Thurston 1743
Escape Rd. near Thurston lava tube, Hawaii Volcanoes National Park....
Escape Rd. near Thurston lava tube, Hawaii Volcanoes National Park. Theridion grallator, happyface spider.

Theridion grallator black Thurston 1741
Escape Rd. near Thurston lava tube, Hawaii Volcanoes National Park....
Escape Rd. near Thurston lava tube, Hawaii Volcanoes National Park. Theridion grallator, happyface spider.

Theridion grallator courting Thurston 0903
Escape Rd. near Thurston lava tube, May 14 2004, at night. This is a...
Escape Rd. near Thurston lava tube, May 14 2004, at night. This is a courting pair on a Clermontia leaf. The male is plucking the web with his third legs.

Theridion grallator courting Thurston 0907
Escape Rd. near Thurston lava tube, May 14 2004, at night. This is a...
Escape Rd. near Thurston lava tube, May 14 2004, at night. This is a courting pair on a Clermontia leaf. The male is plucking the web with his third legs.

Theridion grallator courting Thurston 0908
Escape Rd. near Thurston lava tube, May 14 2004, at night. This is a...
Escape Rd. near Thurston lava tube, May 14 2004, at night. This is a courting pair on a Clermontia leaf.

Escape Rd. near Thurston lava tube, May 14 2004, at night. This is a courting pair on a Clermontia leaf.

Theridion grallator female Thurston 0905
Escape Rd. near Thurston lava tube, May 14 2004, at night. There were quite...
Escape Rd. near Thurston lava tube, May 14 2004, at night. There were quite a few happyface spiders in the area, hanging out under leaves.

Theridion grallator male Thurston 0906
Escape Rd. near Thurston lava tube, May 14 2004, at night. There were quite...
Escape Rd. near Thurston lava tube, May 14 2004, at night. There were quite a few happyface spiders in the area, hanging out under leaves.

Theridion grallator red Thurston 1767
Escape Rd. near Thurston lava tube, Hawaii Volcanoes National Park. Theridion grallator, happyface spider (all red form).

Theridion grallator red Thurston 1768
Escape Rd. near Thurston lava tube, Hawaii Volcanoes National Park. Theridion grallator, happyface spider (all red form).

Theridion grallator red Thurston 1771
Escape Rd. near Thurston lava tube, Hawaii Volcanoes National Park. Theridion grallator, happyface spider (all red form).

Theridion grallator Thurston 0854

Theridion grallator Thurston 0862

Theridion grallator Thurston eating 1847
Escape Rd. near Thurston lava tube, Hawaii Volcanoes National Park. Theridion grallator, happyface spider (all black form).

Theridion grallator Thurston eating 1848
Escape Rd. near Thurston lava tube, Hawaii Volcanoes National Park. Theridion grallator, happyface spider (all black form).

Theridion grallator Thurston eating 1851
Escape Rd. near Thurston lava tube, Hawaii Volcanoes National Park. Theridion grallator, happyface spider (all black form).

Theridion grallator Thurston eggs 1926
Escape Rd. near Thurston lava tube, Hawaii Volcanoes National Park. Theridion grallator, happyface spider. This is a female guarding an egg sac.

Theridion grallator Thurston eggs 1928
Escape Rd. near Thurston lava tube, Hawaii Volcanoes National Park. Theridion grallator, happyface spider. This is a female guarding an egg sac.

Theridion grallator Thurston eggs 1929
Escape Rd. near Thurston lava tube, Hawaii Volcanoes National Park. Theridion grallator, happyface spider. This is a female guarding an egg sac.

Theridion grallator Thurston spiderlings 1966
Escape Rd. near Thurston lava tube, Hawaii Volcanoes National Park. Theridion grallator, happyface spider. Newly hatched spiderlings with the mother.

Theridion grallator Thurston spiderlings 1968
Escape Rd. near Thurston lava tube, Hawaii Volcanoes National Park. Theridion grallator, happyface spider. Newly hatched spiderlings with the mother.

Theridion grallator Thurston spiderlings 1971
Escape Rd. near Thurston lava tube, Hawaii Volcanoes National Park. Theridion grallator, happyface spider. Newly hatched spiderlings with the mother.

Schrankia sp Thurston 5735
July 29, 2010. Thurston lava tube. Schrankia sp., a cave-dwelling...
July 29, 2010. Thurston lava tube. Schrankia sp., a cave-dwelling caterpillar feeding on ohia roots.

Drosophila kambysellisi Lau 0522
July 24, 2009. Laupahoehoe Forest Reserve. Drosophila kambysellisi.

Drosophila kambysellisi Lau 0523
July 24, 2009. Laupahoehoe Forest Reserve. Drosophila kambysellisi.

Drosophila kambysellisi Lau 0525
July 24, 2009. Laupahoehoe Forest Reserve. Drosophila kambysellisi.

Drosophila murphyi Lau 0508
July 24, 2009. Laupahoehoe Forest Reserve. Drosophila murphyi.

Drosophila murphyi Lau 0509
July 24, 2009. Laupahoehoe Forest Reserve. Drosophila murphyi.

Drosophila murphyi Lau 0510
July 24, 2009. Laupahoehoe Forest Reserve. Drosophila murphyi.

Drosophila murphyi Lau 0516a

Megalagrion amaurodytum Kilau 5500
June 9, 2010. Laupahoehoe NAR, above Kilau Gulch. Megalagrion amaurodytum...
June 9, 2010. Laupahoehoe NAR, above Kilau Gulch. Megalagrion amaurodytum peles.

Scotorythra koa defoliation Humuula 9269
Partially defoliation of koa by the native moth Scotorythra paludicola at...
Partially defoliation of koa by the native moth Scotorythra paludicola at about 4000 feet elevation, Humuula Trail. Note the moth flying around the leaves.

Scotorythra koa defoliation Humuula 9272
Defoliation of koa by the native moth Scotorythra paludicola at about 4000...
Defoliation of koa by the native moth Scotorythra paludicola at about 4000 feet elevation, Humuula Trail. The damage is worst at this elevation but stops rather abruptly just above here, suggesting the moths are moving mauka. Adults could also be seen flying near the ground mauka of the defoliated area, but not in damaged areas below about 3800 feet.

Scotorythra paludicola Humuula 9263
Scotorythra paludicola, koa moth. Humuula Trail. This native moth...
Scotorythra paludicola, koa moth. Humuula Trail. This native moth occasionally has outbreaks that defoliate koa trees over huge areas. One of these is currently going on across the eastern slope of Mauna Kea. When these pictures were taken, the current generation of adults was nearing the end of their lives. Hundreds of dead moths were scattered over the ground, and most of the living ones were rather dull-colored due to age.

Scotorythra paludicola Humuula 9264
Scotorythra paludicola, koa moth. Humuula Trail. This native moth...
Scotorythra paludicola, koa moth. Humuula Trail. This native moth occasionally has outbreaks that defoliate koa trees over huge areas. One of these is currently going on across the eastern slope of Mauna Kea. When these pictures were taken, the current generation of adults was nearing the end of their lives. Hundreds of dead moths were scattered over the ground, and most of the living ones were rather dull-colored due to age.

Scotorythra paludicola Humuula 9247
Scotorythra paludicola, koa moth. Humuula Trail. This native moth...
Scotorythra paludicola, koa moth. Humuula Trail. This native moth occasionally has outbreaks that defoliate koa trees over huge areas. One of these is currently going on across the eastern slope of Mauna Kea. When these pictures were taken, the current generation of adults was nearing the end of their lives. Hundreds of dead moths were scattered over the ground, and most of the living ones were rather dull-colored due to age.

Scotorythra paludicola, koa moth caterpillar. Humuula Trail.

Defoliation of a koa seedling by Scotorythra paludicola, koa moth. Humuula...
Defoliation of a koa seedling by Scotorythra paludicola, koa moth. Humuula Trail.

Scotorythra paludicola larva Humuula 3579
Caterpillar of Scotorythra paludicola, koa moth. Large numbers of fallen...
Caterpillar of Scotorythra paludicola, koa moth. Large numbers of fallen caterpillars were visible on the vegetation in this stretch of forest, and with binoculars many could be seen on the phyllodes of koa trees. Frass from the caterpillars covered the ground and leaves.

Scotorythra paludicola larva Humuula 3580
Caterpillar of Scotorythra paludicola, koa moth. Large numbers of fallen...
Caterpillar of Scotorythra paludicola, koa moth. Large numbers of fallen caterpillars were visible on the vegetation in this stretch of forest, and with binoculars many could be seen on the phyllodes of koa trees. Frass from the caterpillars covered the ground and leaves.

Scotorythra paludicola larva Humuula 3581
Caterpillar of Scotorythra paludicola, koa moth. Large numbers of fallen...
Caterpillar of Scotorythra paludicola, koa moth. Large numbers of fallen caterpillars were visible on the vegetation in this stretch of forest, and with binoculars many could be seen on the phyllodes of koa trees. Frass from the caterpillars covered the ground and leaves.

Caterpillars of Scotorythra paludicola, koa moth. Humuula Trail. By this...
Caterpillars of Scotorythra paludicola, koa moth. Humuula Trail. By this time (March 6, 2013), defoliation was occurring at the lower elevation area, around 890m. Large numbers of caterpillars were dropping from the trees and crawling over other plants, sometimes trying to feed on them.

Scotorythra paludicola frass Humuula 3577
Frass from caterpillars of Scotorythra paludicola, koa moth. Large numbers...
Frass from caterpillars of Scotorythra paludicola, koa moth. Large numbers of fallen caterpillars were visible on the vegetation in this stretch of forest, and with binoculars many could be seen on the phyllodes of koa trees. Frass from the caterpillars covered the ground and leaves.

Scotorythra paludicola frass Humuula 3578
Frass from caterpillars of Scotorythra paludicola, koa moth. Large numbers...
Frass from caterpillars of Scotorythra paludicola, koa moth. Large numbers of fallen caterpillars were visible on the vegetation in this stretch of forest, and with binoculars many could be seen on the phyllodes of koa trees. Frass from the caterpillars covered the ground and leaves.

Caterpillar of Scotorythra paludicola, koa moth. Humuula Trail. By this...
Caterpillar of Scotorythra paludicola, koa moth. Humuula Trail. By this time (March 6, 2013), defoliation was occurring at the lower elevation area, around 890m. Large numbers of caterpillars were dropping from the trees and crawling over other plants, sometimes trying to feed on them, like this one on strawberry guava.

Scotorythra paludicola frass Humuula 9355
Caterpillar of Scotorythra paludicola, koa moth. Humuula Trail. By this...
Caterpillar of Scotorythra paludicola, koa moth. Humuula Trail. By this time (March 6, 2013), defoliation was occurring at the lower elevation area, around 890m. Large numbers of caterpillars were dropping from the trees and crawling over other plants, sometimes trying to feed on them.

Frass from caterpillars of Scotorythra paludicola, koa moth. Humuula Trail,...
Frass from caterpillars of Scotorythra paludicola, koa moth. Humuula Trail, 890 m. In this area the frass formed a layer over 1/4" deep, partly due to it being washed down the trail.

Hyposmocoma bugle Kaawalii 9287
Larva of aquatic Hyposmocoma on rocks, Kaawalii stream. These were quite...
Larva of aquatic Hyposmocoma on rocks, Kaawalii stream. These were quite abundant here.

Hyposmocoma bugle Kaawalii 9288
Larva of aquatic Hyposmocoma on rocks, Kaawalii stream. These were quite...
Larva of aquatic Hyposmocoma on rocks, Kaawalii stream. These were quite abundant here.

Hyposmocoma burrito Kaawalii 9283
Larva of aquatic Hyposmocoma on rocks, Kaawalii stream. These were quite...
Larva of aquatic Hyposmocoma on rocks, Kaawalii stream. These were quite abundant here.

Hyposmocoma burrito Kaawalii 9284

Hyposmocoma burrito Kaawalii 9286

Hyposmocoma on rocks Kaawalii 9290

Sophora seed borer 1213
Beetle found feeding in seeds of mamane (Sophora chrysophylla) at Kipuka...
Beetle found feeding in seeds of mamane (Sophora chrysophylla) at Kipuka Nene, Hawaii Volcanoes.

Sophora seed borer 1214
Beetle found feeding in seeds of mamane (Sophora chrysophylla) at Kipuka...
Beetle found feeding in seeds of mamane (Sophora chrysophylla) at Kipuka Nene, Hawaii Volcanoes.

Sophora seed borer 1215
Beetle found feeding in seeds of mamane (Sophora chrysophylla) at Kipuka...
Beetle found feeding in seeds of mamane (Sophora chrysophylla) at Kipuka Nene, Hawaii Volcanoes.

Sophora seed borer 1698 ventral

Sophora seed borer 1703 dorsal

Sophora seed borer 1704 lateral

Sophora seed borer 1705 lateral

Sesbania tomentosa aphids Kipuka Nene 1191
Heavy aphid damage on the endangered ohai, Sesbania tomentosa. Kipuka Nene,...
Heavy aphid damage on the endangered ohai, Sesbania tomentosa. Kipuka Nene, Hawaii Volcanoes.

Sesbania tomentosa aphids Kipuka Nene 1192
Heavy aphid damage on the endangered ohai, Sesbania tomentosa. Kipuka Nene,...
Heavy aphid damage on the endangered ohai, Sesbania tomentosa. Kipuka Nene, Hawaii Volcanoes.

Caconemobius Kahue cave 1317
Caconemobius sp., probably C. varius. Kahue cave.

Caconemobius Kahue cave 1327
Caconemobius sp., probably C. varius. Kahue cave.

Schrankia Kahue cave 1323
Schrankia sp., a moth that lives in caves feeding on the ohia roots that...
Schrankia sp., a moth that lives in caves feeding on the ohia roots that penetrate the ceiling. Kahue cave.

Drosophila hawaiiensis Kapapala
July 2, 2009. Kapapala Canoe Management Area. Drosophila hawaiiensis.

Myoporum dieoff PTA 2541
This kipuka was almost entirely naio, and since 2009 about 90-95% have died....
This kipuka was almost entirely naio, and since 2009 about 90-95% have died. Other sites aren't quite so bad, but have still had heavy losses.

Myoporum thrips damage PTA 2542
Myoporum sandwicense, naio, with heavy damage from naio thrips (Klambothrips...
Myoporum sandwicense, naio, with heavy damage from naio thrips (Klambothrips myopori). Many of the trees in this area were dead from thrips damage.

wood boring lep PTA 2544
Wood-boring moth in Chamaesyce olowaluana. PTA Area 23.

Allograpta obliqua Kipuka Alala 9105
Allograpta obliqua, hover fly, on Myoporum sandwicense. Pohakuloa, Kipuka...
Allograpta obliqua, hover fly, on Myoporum sandwicense. Pohakuloa, Kipuka Alala.

Empicoris rubromaculatus Kipuka Alala 9097
Empicoris rubromaculatus, on Myoporum sandwicense. Pohakuloa, Kipuka Alala.

Klambothrips myopori Kipuka Alala 9091
Klambothrips myopori, naio thrips, on Myoporum sandwicense. Pohakuloa,...
Klambothrips myopori, naio thrips, on Myoporum sandwicense. Pohakuloa, Kipuka Alala.

Klambothrips myopori Kipuka Alala 9094
Klambothrips myopori, naio thrips, on Myoporum sandwicense. Pohakuloa,...
Klambothrips myopori, naio thrips, on Myoporum sandwicense. Pohakuloa, Kipuka Alala.

Nesophrosyne giffardi Kipuka Alala 9099
Nesophrosyne giffardi, on Myoporum sandwicense. Pohakuloa, Kipuka Alala.

Nesophrosyne giffardi Kipuka Alala 9100
Nesophrosyne giffardi, on Myoporum sandwicense. Pohakuloa, Kipuka Alala.

Nesophrosyne sp Kipuka Alala 9111
Nesophrosyne sp., on Myoporum sandwicense. Pohakuloa, Kipuka Alala.

Drosophila ciliaticrus Leilani 6278
Leilani Estates. Drosophila pupa in bark-flux of Tetraplasandra hawaiiensis...
Leilani Estates. Drosophila pupa in bark-flux of Tetraplasandra hawaiiensis (ohe ohe).

Drosophila ciliaticrus Leilani 6281
Leilani Estates. Drosophila pupa in bark-flux of Tetraplasandra hawaiiensis...
Leilani Estates. Drosophila pupa in bark-flux of Tetraplasandra hawaiiensis (ohe ohe).

Plagithmysus Zankau Lupea 2569
Plagithmysus sp. reared from dead wood of Zanthoxylum kauaense from Lupea....
Plagithmysus sp. reared from dead wood of Zanthoxylum kauaense from Lupea. It emerged after only about two weeks. It clearly doesn't match any of the species from Hawaii recorded from Rutaceae or unknown hosts, but they're not always strictly host-specific.

Plagithmysus Zankau Lupea 2570
Plagithmysus sp. reared from dead wood of Zanthoxylum kauaense from Lupea....
Plagithmysus sp. reared from dead wood of Zanthoxylum kauaense from Lupea. It emerged after only about two weeks. It clearly doesn't match any of the species from Hawaii recorded from Rutaceae or unknown hosts, but they're not always strictly host-specific.

Plagithmysus Zankau Lupea 2572
Plagithmysus sp. reared from dead wood of Zanthoxylum kauaense from Lupea....
Plagithmysus sp. reared from dead wood of Zanthoxylum kauaense from Lupea. It emerged after only about two weeks. It clearly doesn't match any of the species from Hawaii recorded from Rutaceae or unknown hosts, but they're not always strictly host-specific.

Ascalapha odorata Lupea 8383
Ascalapha odorata, the black witch. Lupea.

Chrysodeixis eriosoma Lupea 8367
Chrysodeixis eriosoma, garden looper. Lupea.

Chrysodeixis eriosoma Lupea 8368
Chrysodeixis eriosoma, garden looper. Lupea.

Chrysodeixis eriosoma Lupea 8405
Chrysodeixis eriosoma, garden looper. Lupea.

Crambidae Lupea 8379

Crambidae Lupea 8388

Crambidae Lupea 8390

Eudonia Lupea 8364

Eudonia Lupea 8365

Eudonia Lupea 8423

Eupithecia monticolans Lupea 8414

Geometridae Lupea 8366

Geometridae Lupea 8424

Hypocala deflorata Lupea 8409
Hypocala deflorata, Lupea.

Noctuidae Lupea 8359

Noctuidae Lupea 8407

Omiodes Lupea 8430

Scotorythra Lupea 8360

Scotorythra Lupea 8370

Scotorythra Lupea 8377

Udea Lupea 8391

Udea Lupea 8416

Udea Lupea 8420

Drosophila silvestris Kahuku 5966
Kahuku, FBS transect 15, just inside Kau Forest Reserve. Drosophila...
Kahuku, FBS transect 15, just inside Kau Forest Reserve. Drosophila silvestris larvae in rotting Clermontia lindseyana bark.

Drosophila silvestris Kahuku 5968
Kahuku, FBS transect 15, just inside Kau Forest Reserve. Drosophila...
Kahuku, FBS transect 15, just inside Kau Forest Reserve. Drosophila silvestris larvae in rotting Clermontia lindseyana bark.

Drosophila silvestris Kahuku 5956
Kahuku, FBS transect 15, just inside Kau Forest Reserve. Drosophila silvestris.

Kahuku, FBS transect 15, just inside Kau Forest Reserve. Drosophila silvestris.

Trupanea sp Kawainui 7939
Upper Hamakua Ditch Trail, Kawainui. Gall-making Trupanea sp. fruit fly pupa...
Upper Hamakua Ditch Trail, Kawainui. Gall-making Trupanea sp. fruit fly pupa in stem of Dubautia plantaginea, naenae.

Scaptomyza reducta PWW 0777
August 4, 2009. Puu Waawaa Wildlife Sanctuary, 5100', Smilax exclosure....
August 4, 2009. Puu Waawaa Wildlife Sanctuary, 5100', Smilax exclosure. Scaptomyza (Engiscaptomyza) reducta mating on bait sponges. This species was particularly abundant here as well as in South Kona.

Scaptomyza reducta PWW 0781
August 4, 2009. Puu Waawaa Wildlife Sanctuary, 5100', Smilax exclosure....
August 4, 2009. Puu Waawaa Wildlife Sanctuary, 5100', Smilax exclosure. Scaptomyza (Engiscaptomyza) reducta mating on bait sponges. This species was particularly abundant here as well as in South Kona.

Scaptomyza reducta PWW 0784
August 4, 2009. Puu Waawaa Wildlife Sanctuary, 5100', Smilax exclosure....
August 4, 2009. Puu Waawaa Wildlife Sanctuary, 5100', Smilax exclosure. Scaptomyza (Engiscaptomyza) reducta mating on bait sponges. This species was particularly abundant here as well as in South Kona.

Empicoris whitei Kealakekua 8302
Empicoris whitei, thread-legged bug. Kealakekua Ranch.

Vespula pensylvanica nest Kealakekua 8308
Vespula nest, built completely above ground on a fallen log. Kealakekua Ranch.

Vespula pensylvanica nest Kealakekua 8311
Vespula nest, built completely above ground on a fallen log. Kealakekua Ranch.

Vespula pensylvanica nest Kealakekua 8312
Vespula nest, built completely above ground on a fallen log. Kealakekua Ranch.

Drosophila silvestris Kilohana 3027
November 24, 2009. Kilohana exclosure, Puu O Umi NAR. Drosophila silvestris.

Drosophila sproati Kilohana 3054
November 25, 2009. Kilohana exclosure, Puu O Umi NAR. Drosophila sproati.

Drosophila murphyi Kilohana 3028
November 24, 2009. Kilohana exclosure, Puu O Umi NAR. Drosophila murphyi.

Drosophila murphyi Kilohana 3030
November 24, 2009. Kilohana exclosure, Puu O Umi NAR. Drosophila murphyi.

Drosophila murphyi Kilohana 3032
November 24, 2009. Kilohana exclosure, Puu O Umi NAR. Drosophila murphyi.

Drosophila tanythrix Kilohana 0707
August 3, 2009. Kilohana exclosure, Puu O Umi NAR. Drosophila tanythrix...
August 3, 2009. Kilohana exclosure, Puu O Umi NAR. Drosophila tanythrix courtship.

Drosophila tanythrix Kilohana 0708
August 3, 2009. Kilohana exclosure, Puu O Umi NAR. Drosophila tanythrix...
August 3, 2009. Kilohana exclosure, Puu O Umi NAR. Drosophila tanythrix courtship.

Drosophila ochrobasis Kilohana 3115
November 25, 2009. Kilohana exclosure, Puu O Umi NAR. Drosophila ochrobasis.

Drosophila ochrobasis Kilohana 3116
November 25, 2009. Kilohana exclosure, Puu O Umi NAR. Drosophila ochrobasis.

Scaptomyza undulata Kilohana 0676
August 3, 2009. Kilohana exclosure, Puu O Umi NAR. Scaptomyza...
August 3, 2009. Kilohana exclosure, Puu O Umi NAR. Scaptomyza (Grimshawomyia) undulata on Astelia menziesii.

Scaptomyza undulata Kilohana 0677
August 3, 2009. Kilohana exclosure, Puu O Umi NAR. Scaptomyza...
August 3, 2009. Kilohana exclosure, Puu O Umi NAR. Scaptomyza (Grimshawomyia) undulata on Astelia menziesii.

Scaptomyza undulata Kilohana 0682
August 3, 2009. Kilohana exclosure, Puu O Umi NAR. Scaptomyza...
August 3, 2009. Kilohana exclosure, Puu O Umi NAR. Scaptomyza (Grimshawomyia) undulata on Astelia menziesii.

Drosophila ochrobasis Kilohana 3122
November 25, 2009. Kilohana exclosure, Puu O Umi NAR. Drosophila ochrobasis.

Drosophila silvestris Kilohana 5150
May 24, 2010. Kilohana exclosure, Puu O Umi NAR. Drosophila silvestris.

Drosophila silvestris Kilohana 5154
May 24, 2010. Kilohana exclosure, Puu O Umi NAR. Drosophila silvestris.

May 24, 2010. Kilohana exclosure, Puu O Umi NAR. Drosophila silvestris.

Drosophila sp dance Kilohana 5298
May 25, 2010. Kilohana exclosure, Puu O Umi NAR. These two male Drosophila...
May 25, 2010. Kilohana exclosure, Puu O Umi NAR. These two male Drosophila (undescribed ciliated tarsus species) engaged in this circling dance for at least five minutes until I scared them off.

Drosophila sp dance Kilohana 5300
May 25, 2010. Kilohana exclosure, Puu O Umi NAR. These two male Drosophila...
May 25, 2010. Kilohana exclosure, Puu O Umi NAR. These two male Drosophila (undescribed ciliated tarsus species) engaged in this circling dance for at least five minutes until I scared them off.

Drosophila sp dance Kilohana 5301
May 25, 2010. Kilohana exclosure, Puu O Umi NAR. These two male Drosophila...
May 25, 2010. Kilohana exclosure, Puu O Umi NAR. These two male Drosophila (undescribed ciliated tarsus species) engaged in this circling dance for at least five minutes until I scared them off.

August 3, 2009. Kilohana exclosure, Puu O Umi NAR. Drosophila silvestris.

August 3, 2009. Kilohana exclosure, Puu O Umi NAR. Drosophila silvestris...
August 3, 2009. Kilohana exclosure, Puu O Umi NAR. Drosophila silvestris and D. sproati.

May 25, 2010. Kilohana exclosure, Puu O Umi NAR. Drosophila ochrobasis (male).

Drosophila sproati Kilohana 5324
May 25, 2010. Kilohana exclosure, Puu O Umi NAR. Drosophila sproati.

Drosophila sproati Kilohana 5328
May 25, 2010. Kilohana exclosure, Puu O Umi NAR. Drosophila sproati.

May 25, 2010. Kilohana exclosure, Puu O Umi NAR. Drosophila sproati.

Drosophila ochrobasis Kilohana1
May 22, 2009. Puu O Umi NAR, Kilohana exclosure. Drosophila ochrobasis male...
May 22, 2009. Puu O Umi NAR, Kilohana exclosure. Drosophila ochrobasis male on bait sponge.

Drosophila ochrobasis Kilohana2
May 22, 2009. Puu O Umi NAR, Kilohana exclosure. Drosophila ochrobasis male...
May 22, 2009. Puu O Umi NAR, Kilohana exclosure. Drosophila ochrobasis male on bait sponge.

Drosophila ochrobasis Kilohana3
May 22, 2009. Puu O Umi NAR, Kilohana exclosure. Drosophila ochrobasis male...
May 22, 2009. Puu O Umi NAR, Kilohana exclosure. Drosophila ochrobasis male on bait sponge.

Drosophila silvestris Kilohana
May 22, 2009. Puu O Umi NAR, Kilohana exclosure. Drosophila silvestris on...
May 22, 2009. Puu O Umi NAR, Kilohana exclosure. Drosophila silvestris on bait sponge.

Drosophila sproati Kilohana
May 22, 2009. Puu O Umi NAR, Kilohana exclosure. Drosophila sproati on bait...
May 22, 2009. Puu O Umi NAR, Kilohana exclosure. Drosophila sproati on bait sponge.

November 25, 2009. Kilohana exclosure, Puu O Umi NAR. Drosophila sproati.

Megalagrion hawaiiense Kilohana 5213
May 24, 2010. Kilohana exclosure, Puu O Umi NAR. Megalagrion hawaiiense.

Megalagrion hawaiiense Kilohana 5215
May 24, 2010. Kilohana exclosure, Puu O Umi NAR. Megalagrion hawaiiense.

Megalagrion hawaiiense Kilohana 5225
May 24, 2010. Kilohana exclosure, Puu O Umi NAR. Megalagrion hawaiiense.

Drosophila heteroneura Kukuiopae 7876
South Kona Forest Reserve, Kukuiopae section, 4900'. Drosophila heteroneura...
South Kona Forest Reserve, Kukuiopae section, 4900'. Drosophila heteroneura female. The head is broader than normal for Drosophila but less so than the males.

Drosophila heteroneura Kukuiopae 7881
South Kona Forest Reserve, Kukuiopae section, 4900'. Drosophila heteroneura...
South Kona Forest Reserve, Kukuiopae section, 4900'. Drosophila heteroneura female. The head is broader than normal for Drosophila but less so than the males.

Drosophila heteroneura Kukuiopae 7886
South Kona Forest Reserve, Kukuiopae section, 4900'. Drosophila heteroneura...
South Kona Forest Reserve, Kukuiopae section, 4900'. Drosophila heteroneura female. The head is broader than normal for Drosophila but less so than the males.

Drosophila silvestris Kukuiopae 7870
South Kona Forest Reserve, Kukuiopae section, 4900'. Drosophila silvestris.

Drosophila silvestris Kukuiopae 7871
South Kona Forest Reserve, Kukuiopae section, 4900'. Drosophila silvestris.

Drosophila silvestris Kukuiopae 7894
South Kona Forest Reserve, Kukuiopae section, 4900'. Drosophila silvestris.

Hylaeus anthracinus Puako 0533
Puako beach. Hylaeus anthracinus on tree heliotrope, Tournefortia argentea....
Puako beach. Hylaeus anthracinus on tree heliotrope, Tournefortia argentea. This is the only exotic plant that the native bees regularly visit.

Hylaeus anthracinus Puako 0534
Puako beach. Hylaeus anthracinus on tree heliotrope, Tournefortia argentea....
Puako beach. Hylaeus anthracinus on tree heliotrope, Tournefortia argentea. This is the only exotic plant that the native bees regularly visit.

Hylaeus anthracinus Puako 0535
Puako beach. Hylaeus anthracinus on tree heliotrope, Tournefortia argentea....
Puako beach. Hylaeus anthracinus on tree heliotrope, Tournefortia argentea. This is the only exotic plant that the native bees regularly visit.

Hylaeus anthracinus habitat Puako 3649
Typical habitat for Hylaeus anthracinus. Puako, South Kohala. Coral rubble...
Typical habitat for Hylaeus anthracinus. Puako, South Kohala. Coral rubble for nesting is abundant and the bees visit flowers of tree heliotropes (Heliotropium foertherianum, =Tournefortia argentea) in large numbers. There was over 100 bees in the frame when this picture was taken.

Hylaeus anthracinus habitat Puako 3652
Typical habitat for Hylaeus anthracinus. Puako, South Kohala. Coral rubble...
Typical habitat for Hylaeus anthracinus. Puako, South Kohala. Coral rubble for nesting is abundant and the bees visit flowers of tree heliotropes (Heliotropium foertherianum, =Tournefortia argentea) in large numbers. There was over 100 bees in the frame when this picture was taken.

Myoporum sandwicense naio thrips MKBH 2650
Myoporum sandwicense hit by naio thrips north of the Mauna Kea Beach Hotel.

Myoporum sandwicense naio thrips MKBH 2651
Myoporum sandwicense hit by naio thrips north of the Mauna Kea Beach Hotel.

Drosophila fungiperda Kahuku 7248
Kahuku, Hawaii Volcanoes NP. Drosophila fungiperda. Males lekking on ohia...
Kahuku, Hawaii Volcanoes NP. Drosophila fungiperda. Males lekking on ohia leaves would fight each other if another landed on their leaf.

Drosophila fungiperda Kahuku 7253
Kahuku, Hawaii Volcanoes NP. Drosophila fungiperda. Males lekking on ohia...
Kahuku, Hawaii Volcanoes NP. Drosophila fungiperda. Males lekking on ohia leaves would fight each other if another landed on their leaf.

Drosophila fungiperda Kahuku 7254
Kahuku, Hawaii Volcanoes NP. Drosophila fungiperda. Males lekking on ohia...
Kahuku, Hawaii Volcanoes NP. Drosophila fungiperda. Males lekking on ohia leaves would fight each other if another landed on their leaf.

Drosophila conspicua Kukuiopae 3438
December 29, 2009. South Kona Forest Reserve, Kukuiopae section, 4600'....
December 29, 2009. South Kona Forest Reserve, Kukuiopae section, 4600'. Drosophila conspicua, male.

Drosophila conspicua Kukuiopae 3477
December 29, 2009. South Kona Forest Reserve, Kukuiopae section, 4600'....
December 29, 2009. South Kona Forest Reserve, Kukuiopae section, 4600'. Drosophila conspicua, female.

Drosophila conspicua Kukuiopae 3494
December 29, 2009. South Kona Forest Reserve, Kukuiopae section, 4600'....
December 29, 2009. South Kona Forest Reserve, Kukuiopae section, 4600'. Drosophila conspicua, female.

Drosophila formella Kukuiopae 3425
December 29, 2009. South Kona Forest Reserve, Kukuiopae section, 4600'. Drosophila formella.

Drosophila formella Kukuiopae 3428
December 29, 2009. South Kona Forest Reserve, Kukuiopae section, 4600'. Drosophila formella.

Drosophila formella Kukuiopae 3430
December 29, 2009. South Kona Forest Reserve, Kukuiopae section, 4600'. Drosophila formella.

December 29, 2009. South Kona Forest Reserve, Kukuiopae section, 4600'. Drosophila heteroneura. This was the first sighting of the species in at least eight years, and the first time recently it had been recorded from Kukuiopae.

Drosophila heteroneura Kukuiopae 3490b

Drosophila on sponge Kukuiopae 3445
December 29, 2009. South Kona Forest Reserve, Kukuiopae section, 4600'. Drosophila bait sponge with D. heteroneura, D. silvestris, D. murphyi, D. hawaiiensis, D. sproati, and D. tanythrix. Drosophila conspicua and D. formella were also at this site.

December 29, 2009. South Kona Forest Reserve, Kukuiopae section, 4600'. Drosophila silvestris.

Erythrina variegata galled Waikoloa 2643
Erythrina variegata? This was out in the middle of the anchialine pond area...
Erythrina variegata? This was out in the middle of the anchialine pond area at the Hilton Waikoloa and was hit much harder by gall wasps than any other Erythrina I've seen in a long time, especially on the twigs.

Erythrina variegata galled Waikoloa 2644
Erythrina variegata? This was out in the middle of the anchialine pond area...
Erythrina variegata? This was out in the middle of the anchialine pond area at the Hilton Waikoloa and was hit much harder by gall wasps than any other Erythrina I've seen in a long time, especially on the twigs.

South Kona Forest Reserve, Kukuiopae. Drosophila conspicua.

South Kona Forest Reserve, Kukuiopae. Drosophila conspicua, with D. murphyi...
South Kona Forest Reserve, Kukuiopae. Drosophila conspicua, with D. murphyi (below left and center) and alien D. immigrans (below right).

South Kona Forest Reserve, Kukuiopae. Drosophila conspicua, with alien D....
South Kona Forest Reserve, Kukuiopae. Drosophila conspicua, with alien D. immigrans (right).

August 5, 2009. South Kona Forest Reserve, Kukuiopae section, 3750'....
August 5, 2009. South Kona Forest Reserve, Kukuiopae section, 3750'. Drosophila formella.

Drosophila murphyi Kukuiopae 0860
August 5, 2009. South Kona Forest Reserve, Kukuiopae section, 3750'....
August 5, 2009. South Kona Forest Reserve, Kukuiopae section, 3750'. Drosophila murphyi.

Drosophila murphyi Kukuiopae 0861
August 5, 2009. South Kona Forest Reserve, Kukuiopae section, 3750'....
August 5, 2009. South Kona Forest Reserve, Kukuiopae section, 3750'. Drosophila murphyi.

Drosophila prolaticilia Kukuiopae 0872
August 5, 2009. South Kona Forest Reserve, Kukuiopae section, 3750'. Drosophila prolaticilia.

Drosophila prolaticilia Kukuiopae 0873
August 5, 2009. South Kona Forest Reserve, Kukuiopae section, 3750'. Drosophila prolaticilia.

Drosophila prolaticilia Kukuiopae 0874
August 5, 2009. South Kona Forest Reserve, Kukuiopae section, 3750'. Drosophila prolaticilia.

Ceratina dentipes Puho 0975
Ceratina dentipes, small carpenter bee (introduced). Puuhonua O Honaunau NHP.

Ceratina dentipes Puho 0977
Ceratina dentipes, small carpenter bee (introduced). Puuhonua O Honaunau NHP.

Hylaeus anthracinus Puho 0971
Hylaeus anthracinus, Puuhonua O Honaunau NHP.

Hylaeus anthracinus Puho 0974
Hylaeus anthracinus, Puuhonua O Honaunau NHP.

Hylaeus anthracinus Puho 0999
Hylaeus anthracinus nest in coral rock, Puuhonua O Honaunau NHP. While bees...
Hylaeus anthracinus nest in coral rock, Puuhonua O Honaunau NHP. While bees have often been seen entering holes in coral rubble and seem to be most abundant where there is a lot of it, this is the first definite evidence that they're nesting in it. The cellophane-like substance is a secretion that the female bee produces to line the nest cell and make it waterproof; when filled up, she seals it up like this until the new adults emerge.

Hylaeus anthracinus Puho 0999a
Hylaeus anthracinus nest in coral rock, Puuhonua O Honaunau NHP. While bees...
Hylaeus anthracinus nest in coral rock, Puuhonua O Honaunau NHP. While bees have often been seen entering holes in coral rubble and seem to be most abundant where there is a lot of it, this is the first definite evidence that they're nesting in it. The cellophane-like substance is a secretion that the female bee produces to line the nest cell and make it waterproof; when filled up, she seals it up like this until the new adults emerge.

Hylaeus anthracinus, Puuhonua O Honaunau NHP.

Hylaeus anthracinus Puho 1006a

Ceratina dentipes Puho 0995
Ceratina dentipes, small carpenter bee (introduced). Puuhonua O Honaunau NHP.

Hylaeus anthracinus Puho 0991a

Puho reintro site 1 4593
PUHO Hylaeus anthracinus reintroduction site 1, at Alahaka Bay near where the...
PUHO Hylaeus anthracinus reintroduction site 1, at Alahaka Bay near where the coastal trail meets the inland 1871 trail. There is a high diversity of native coastal plants here, including ilima, ohai, naupaka, uhaloa, pau o Hiiaka, maiapilo, milo, and kou, as well as tree heliotrope.

Amphicerus cornutus PUHO 4589
Amphicerus cornutus, an introduced bostrichid beetle, boring into twigs of...
Amphicerus cornutus, an introduced bostrichid beetle, boring into twigs of naupaka. Puuhonua O Honaunau NHP.

Hylaeus anthracinus PUHO 4591
Hylaeus anthracinus, successfully reintroduced at Puuhonua O Honaunau NHP.

Hylaeus anthracinus PUHO 4592
Hylaeus anthracinus, successfully reintroduced at Puuhonua O Honaunau NHP.

Hylaeus attacked by Pheidole PUHO 6312
North of Alahaka Bay, reintroduction of Hylaeus was attempted at Puuhonua O...
North of Alahaka Bay, reintroduction of Hylaeus was attempted at Puuhonua O Honaunau NHP. However, large populations of big-headed ants (Pheidole megacephala) are present. Within seconds of bees being released, ants attacked the bees that were still unable to fly from being transported on ice.

Hylaeus attacked by Pheidole PUHO 6313
North of Alahaka Bay, reintroduction of Hylaeus was attempted at Puuhonua O...
North of Alahaka Bay, reintroduction of Hylaeus was attempted at Puuhonua O Honaunau NHP. However, large populations of big-headed ants (Pheidole megacephala) are present. Within seconds of bees being released, ants attacked the bees that were still unable to fly from being transported on ice.

Hylaeus attacked by Pheidole PUHO 6314
North of Alahaka Bay, reintroduction of Hylaeus was attempted at Puuhonua O...
North of Alahaka Bay, reintroduction of Hylaeus was attempted at Puuhonua O Honaunau NHP. However, large populations of big-headed ants (Pheidole megacephala) are present. Within seconds of bees being released, ants attacked the bees that were still unable to fly from being transported on ice.

Puho reintro site 2 4596
PUHO Hylaeus anthracinus reintroduction site 2, near the beginning of the...
PUHO Hylaeus anthracinus reintroduction site 2, near the beginning of the coastal trail. There are Sesbania tomentosa, naupaka, and lots of Waltheria coming up here.

Ethmia nigroapicella damage Honaunau 3655
Serious defoliation of kou (Cordia subcordata) by the kou leafworm, Ethmia...
Serious defoliation of kou (Cordia subcordata) by the kou leafworm, Ethmia nigroapicella. Puuhonua o Honaunau NHP. The closest tree is resprouting from the base.

Ethmia nigroapicella damage Honaunau 3656
Serious defoliation of kou (Cordia subcordata) by the kou leafworm, Ethmia...
Serious defoliation of kou (Cordia subcordata) by the kou leafworm, Ethmia nigroapicella. Puuhonua o Honaunau NHP. The closest tree is resprouting from the base.

Ethmia nigroapicella Honaunau 3653
Ethmia nigroapicella, kou leafworm, on a shoot of kou (Cordia subcordata)....
Ethmia nigroapicella, kou leafworm, on a shoot of kou (Cordia subcordata). Puuhonua o Honaunau NHP.

Vespula nest Kukuiopae 0856
August 5, 2009. South Kona Forest Reserve, Kukuiopae section, 2300'. An...
August 5, 2009. South Kona Forest Reserve, Kukuiopae section, 2300'. An enormous Vespula nest, built along the side of, into, and under a big ohia tree just below where the road enters the lava flow.

Vespula nest Kukuiopae 0854
August 5, 2009. South Kona Forest Reserve, Kukuiopae section, 2300'. An...
August 5, 2009. South Kona Forest Reserve, Kukuiopae section, 2300'. An enormous Vespula nest, built along the side of, into, and under a big ohia tree just below where the road enters the lava flow.

Vespula nest Kukuiopae 3497
December 29, 2009. South Kona Forest Reserve, Kukuiopae section, 4600'. The...
December 29, 2009. South Kona Forest Reserve, Kukuiopae section, 4600'. The small gully where the Drosophila were collected. Other areas nearby did not have as many, though D. heteroneura was collected elsewhere, indicating it's spread throughout the forest. The entire area is heavily impacted by feral cattle.

December 29, 2009. South Kona Forest Reserve, Kukuiopae section, 4600'. The...
December 29, 2009. South Kona Forest Reserve, Kukuiopae section, 4600'. The small gully where the Drosophila were collected. Other areas nearby did not have as many, though D. heteroneura was collected elsewhere, indicating it's spread throughout the forest. The entire area is heavily impacted by feral cattle.

Hylaeus anthracinus Kiholo 2053
Kiholo, dune area between the Kaupulehu lava flows. Hylaeus anthracinus...
Kiholo, dune area between the Kaupulehu lava flows. Hylaeus anthracinus trapped in a flower of Ipomoea pes-caprae, pohuehue. The bees apparently visited the flowers for pollen and nectar, and were unable to climb up the slippery walls. Alien Ceratina smaragdula bees were also found in some, but they appeared catatonic and weren't trying to get out like the Hylaeus.

Hylaeus anthracinus Kiholo 2055
Kiholo, dune area between the Kaupulehu lava flows. Hylaeus anthracinus...
Kiholo, dune area between the Kaupulehu lava flows. Hylaeus anthracinus trapped in a flower of Ipomoea pes-caprae, pohuehue. The bees apparently visited the flowers for pollen and nectar, and were unable to climb up the slippery walls. Alien Ceratina smaragdula bees were also found in some, but they appeared catatonic and weren't trying to get out like the Hylaeus.

Hylaeus anthracinus Kiholo 2058
Kiholo, dune area between the Kaupulehu lava flows. Hylaeus anthracinus...
Kiholo, dune area between the Kaupulehu lava flows. Hylaeus anthracinus trapped in a flower of Ipomoea pes-caprae, pohuehue. The bees apparently visited the flowers for pollen and nectar, and were unable to climb up the slippery walls. Alien Ceratina smaragdula bees were also found in some, but they appeared catatonic and weren't trying to get out like the Hylaeus.

Hylaeus anthracinus Kohanaiki 2131
Hylaeus anthracinus on maiapilo (Capparis sandwichiana). Kohanaiki.

Drosophila digressa Manuka 0982
August 7, 2009. Manuka NAR, olopua exclosure. Drosophila digressa?...
August 7, 2009. Manuka NAR, olopua exclosure. Drosophila digressa? Coloration is slightly different from those seen in Olaa: the median stripe is fainter, and an anterior median spot is present in the wing.

Drosophila digressa Manuka 0983
August 7, 2009. Manuka NAR, olopua exclosure. Drosophila digressa?...
August 7, 2009. Manuka NAR, olopua exclosure. Drosophila digressa? Coloration is slightly different from those seen in Olaa: the median stripe is fainter, and an anterior median spot is present in the wing. This shows how many were at this spot (9 are present just in this shot), compared to how rare they are at Olaa (~3 specimens in the last 10 years of relatively intensive surveys).

Drosophila digressa Manuka 0978
August 7, 2009. Manuka NAR, olopua exclosure. Drosophila digressa?...
August 7, 2009. Manuka NAR, olopua exclosure. Drosophila digressa? Coloration is slightly different from those seen in Olaa: the median stripe is fainter, and an anterior median spot is present in the wing.

Drosophila spp Manuka 0975
August 7, 2009. Manuka NAR, olopua exclosure. Drosophila digressa? Coloration is slightly different from those seen in Olaa: the median stripe is fainter, and an anterior median spot is present in the wing. This shows how many were at this spot (3 D. digressa plus one D. nr. obatai just in this shot), compared to how rare they are at Olaa (~3 specimens in the last 10 years of relatively intensive surveys).

August 7, 2009. Manuka NAR, olopua exclosure. Drosophila digressa? Coloration is slightly different from those seen in Olaa: the median stripe is fainter, and an anterior median spot is present in the wing.

Drosophila ciliaticrus Manuka 0974
August 7, 2009. Manuka NAR, olopua exclosure. Drosophila ciliaticrus.

Hylaeus difficilis Kamilo 2640
Hylaeus difficilis mating pair in an ilima flower, Kamilo. Naturally, when I...
Hylaeus difficilis mating pair in an ilima flower, Kamilo. Naturally, when I came back with my good camera there weren't even single bees around.

Drosophila cyrtoloma Hana 1276
Hana Forest Reserve, near Boot Camp on the rim of Waihoi Valley. Drosophila...
Hana Forest Reserve, near Boot Camp on the rim of Waihoi Valley. Drosophila cyrtoloma.

Drosophila cyrtoloma Hana 1285
Hana Forest Reserve, near Boot Camp on the rim of Waihoi Valley. Drosophila...
Hana Forest Reserve, near Boot Camp on the rim of Waihoi Valley. Drosophila cyrtoloma.

Drosophila cyrtoloma Hana 1286
Hana Forest Reserve, near Boot Camp on the rim of Waihoi Valley. Drosophila...
Hana Forest Reserve, near Boot Camp on the rim of Waihoi Valley. Drosophila cyrtoloma.

Hana Forest Reserve, near Boot Camp on the rim of Waihoi Valley. Drosophila cyrtoloma, two males about to fight for a lek spot.

Hana Forest Reserve, near Boot Camp on the rim of Waihoi Valley. Drosophila cyrtoloma. The male on the right is attempting court by taking the head-under-wing position and vibrating his wings, but the other fly is also a male.

One fly against the world.

Megalagrion sp Hana 1226
Hana Forest Reserve, near Boot Camp on the rim of Waihoi Valley. A pair of...
Hana Forest Reserve, near Boot Camp on the rim of Waihoi Valley. A pair of Megalagrion damselflies mating.

Trupanea waikamoi Hana 1584
Hana Forest Reserve, below "the crater" at the head of Waihoi Valley....
Hana Forest Reserve, below "the crater" at the head of Waihoi Valley. Trupanea n. sp. "waikamoi".

Drosophila petalopeza Paliku 1941
August 2, 2007. Haleakala NP, Paliku. Large numbers of Drosophila...
August 2, 2007. Haleakala NP, Paliku. Large numbers of Drosophila petalopeza were lekking under leaves near the ranger cabin.

Drosophila petalopeza Paliku 1942
August 2, 2007. Haleakala NP, Paliku. Large numbers of Drosophila...
August 2, 2007. Haleakala NP, Paliku. Large numbers of Drosophila petalopeza were lekking under leaves near the ranger cabin.

Drosophila petalopeza Paliku 2010
August 4, 2007. Haleakala NP, Paliku. Large numbers of Drosophila...
August 4, 2007. Haleakala NP, Paliku. Large numbers of Drosophila petalopeza were lekking under leaves near the ranger cabin.

August 1, 2007. Haleakala NP, Paliku. Drosophila petalopeza males fighting...
August 1, 2007. Haleakala NP, Paliku. Drosophila petalopeza males fighting over a lekking spot on the underside of a Cheirodendron leaf.

August 1, 2007. Haleakala NP, Paliku. Drosophila petalopeza were lekking...
August 1, 2007. Haleakala NP, Paliku. Drosophila petalopeza were lekking under Cheirodendron leaves near the ranger cabin.

Eupithecia sp Paliku 1906
August 1, 2007. Haleakala NP, Paliku. Eupithecia resting on Osteomeles.

Hylaeus difficilis Paliku 2018
August 5, 2007. Haleakala NP, Paliku. Hylaeus difficilis on Oenothera...
August 5, 2007. Haleakala NP, Paliku. Hylaeus difficilis on Oenothera flower, apparently drinking water rather than nectar. Note that the wings are matted together by the rain.

Hylaeus difficilis Paliku 2019
August 5, 2007. Haleakala NP, Paliku. Hylaeus difficilis on Oenothera...
August 5, 2007. Haleakala NP, Paliku. Hylaeus difficilis on Oenothera flower, apparently drinking water rather than nectar. Note that the wings are matted together by the rain.

Hylaeus on silversword 1892
August 1, 2007. Haleakala NP, Halemauu Trail. Hylaeus visiting silversword...
August 1, 2007. Haleakala NP, Halemauu Trail. Hylaeus visiting silversword flower.

Hylaeus on silversword dead 1903
August 1, 2007. Haleakala NP, Halemauu Trail. Dead Hylaeus trapped in the...
August 1, 2007. Haleakala NP, Halemauu Trail. Dead Hylaeus trapped in the sticky hairs on a silversword.

caterpillar Auwahi 1135
Lower Auwahi. Caterpillar (Nymphalidae?) on Nicotiana glauca.

Hylaeus on Dubautia menziesii 2043
August 5, 2007. Haleakala Crater. Hylaeus sp. visiting flower of Dubautia...
August 5, 2007. Haleakala Crater. Hylaeus sp. visiting flower of Dubautia menziesii.

Drosophila oreas Puaakaa 6897
Puaakaa Stream. Drosophila oreas.

Drosophila oreas Puaakaa 6898
Puaakaa Stream. Drosophila oreas.

Drosophila oreas Puaakaa 6901
Puaakaa Stream. Drosophila oreas.

Megalagrion blackburni Puaakaa 6879
Puaakaa Stream. Megalagrion blackburni in tandem.

Megalagrion blackburni Puaakaa 6880
Puaakaa Stream. Megalagrion blackburni in tandem.

Megalagrion nigrohamatum Puaakaa 6888
Puaakaa Stream. Megalagrion nigrohamatum.

Megalagrion nigrohamatum Puaakaa 6891
Puaakaa Stream. Megalagrion nigrohamatum.

Megalagrion nigrohamatum Puaakaa 6894
Puaakaa Stream. Megalagrion nigrohamatum.

Megalagrion sp Puaakaa 6906
Puaakaa Stream. Megalagrion sp.

Hylaeus nivicola Halemauu 2059
August 5, 2007. Haleakala NP, Halemauu Trail above the crater. Hylaeus...
August 5, 2007. Haleakala NP, Halemauu Trail above the crater. Hylaeus nivicola on flowers of Geranium cuneatum.

Hylaeus nivicola Halemauu 2060
August 5, 2007. Haleakala NP, Halemauu Trail above the crater. Hylaeus...
August 5, 2007. Haleakala NP, Halemauu Trail above the crater. Hylaeus nivicola on flowers of Geranium cuneatum.

Hylaeus nivicola Halemauu 2061
August 5, 2007. Haleakala NP, Halemauu Trail above the crater. Hylaeus...
August 5, 2007. Haleakala NP, Halemauu Trail above the crater. Hylaeus nivicola on flowers of Geranium cuneatum.

Megalagrion blackburni E Wailua Iki 0782
East Wailua Iki gulch, just above Hana Highway. Megalagrion blackburni.

Drosophila cyrtoloma Waikamoi1
June 2, 2009. Waikamoi Preserve, unnamed trail paralleling western pali of...
June 2, 2009. Waikamoi Preserve, unnamed trail paralleling western pali of Koolau Gap. Drosophila cyrtoloma on a bait sponge, at around 6200' elevation.

Drosophila cyrtoloma Waikamoi2
June 2, 2009. Waikamoi Preserve, unnamed trail paralleling western pali of...
June 2, 2009. Waikamoi Preserve, unnamed trail paralleling western pali of Koolau Gap. Drosophila cyrtoloma on a bait sponge, at around 6200' elevation.

Megalagrion hawaiiense Waiakamilo 0791
Koolau Ditch trail, Waiokamilo Stream. Megalagrion hawaiiense?

Megalagrion hawaiiense Waiakamilo 0792
Koolau Ditch trail, Waiokamilo Stream. Megalagrion hawaiiense?

Drosophila neopicta Waikamoi1
June 3, 2009. Waikamoi Preserve, transect 3 (parallel to boardwalk)....
June 3, 2009. Waikamoi Preserve, transect 3 (parallel to boardwalk). Drosophila neopicta on bait sponge.

Drosophila neopicta Waikamoi2
June 3, 2009. Waikamoi Preserve, transect 3 (parallel to boardwalk)....
June 3, 2009. Waikamoi Preserve, transect 3 (parallel to boardwalk). Drosophila neopicta on bait sponge.

Drosophila neopicta Waikamoi3
June 3, 2009. Waikamoi Preserve, transect 3 (parallel to boardwalk)....
June 3, 2009. Waikamoi Preserve, transect 3 (parallel to boardwalk). Drosophila neopicta on bait sponge.

Drosophila hamifera Waikamoi 6988
Waikamoi Flume. Drosophila hamifera.

Drosophila hamifera Waikamoi 6989
Waikamoi Flume. Drosophila hamifera.

Drosophila hamifera Waikamoi 6990
Waikamoi Flume. Drosophila hamifera.

Drosophila fasciculisetae Waikamoi 7011
Waikamoi, along preserve boundary fence. Drosophila fasciculisetae.

Drosophila fasciculisetae Waikamoi 7015
Waikamoi, along preserve boundary fence. Drosophila fasciculisetae.

Drosophila fasciculisetae Waikamoi 7016
Waikamoi, along preserve boundary fence. Drosophila fasciculisetae.

Drosophila grimshawi Waikamoi 7019
Waikamoi, along preserve boundary fence. Drosophila grimshawi.

Drosophila grimshawi Waikamoi 7020
Waikamoi, along preserve boundary fence. Drosophila grimshawi.

Drosophila grimshawi Waikamoi 7025
Waikamoi, along preserve boundary fence. Drosophila grimshawi.

Drosophila adiastola Waikamoi 7039
Waikamoi, Haiku Uka, Kula pipeline road. Drosophila adiastola.

Drosophila adiastola Waikamoi 7040
Waikamoi, Haiku Uka, Kula pipeline road. Drosophila adiastola.

Drosophila adiastola Waikamoi 7041
Waikamoi, Haiku Uka, Kula pipeline road. Drosophila adiastola.

Drosophila adunca Waikamoi 7034
Waikamoi, Haiku Uka, Kula pipeline road. Drosophila adunca.

Drosophila clavisetae Waikamoi 1164
Waikamoi, Haiku Uka, Kula Pipeline road. Drosophila clavisetae on rotting...
Waikamoi, Haiku Uka, Kula Pipeline road. Drosophila clavisetae on rotting stem of Cyanea mceldowneyi. Below are two D. adiastola.

Drosophila clavisetae Waikamoi 1165
Waikamoi, Haiku Uka, Kula Pipeline road. Drosophila clavisetae on rotting...
Waikamoi, Haiku Uka, Kula Pipeline road. Drosophila clavisetae on rotting stem of Cyanea mceldowneyi. Below are two D. adiastola.

Drosophila clavisetae Waikamoi 1203
August 20, 2009. Waikamoi (Haiku Uka), Pipeline Road, 3900'. Drosophila...
August 20, 2009. Waikamoi (Haiku Uka), Pipeline Road, 3900'. Drosophila clavisetae.

August 20, 2009. Waikamoi (Haiku Uka), Pipeline Road, 3900'. Drosophila...
August 20, 2009. Waikamoi (Haiku Uka), Pipeline Road, 3900'. Drosophila clavisetae.

Drosophila cyrtoloma Waikamoi 1289
August 20, 2009. Waikamoi (Haiku Uka), Pipeline Road, 3900'. Drosophila cyrtoloma.

Drosophila cyrtoloma Waikamoi 1290
August 20, 2009. Waikamoi (Haiku Uka), Pipeline Road, 3900'. Drosophila cyrtoloma.

Drosophila cyrtoloma Waikamoi 1291
August 20, 2009. Waikamoi (Haiku Uka), Pipeline Road, 3900'. Drosophila cyrtoloma.

Drosophila spp Waikamoi 1170
Waikamoi, Haiku Uka, Kula Pipeline road. Drosophila clavisetae (left) and D....
Waikamoi, Haiku Uka, Kula Pipeline road. Drosophila clavisetae (left) and D. truncipenna (right) on rotting stem of Cyanea mceldowneyi.

Drosophila truncipenna Waikamoi 7038
Waikamoi, Haiku Uka, Kula pipeline road. Drosophila truncipenna, male...
Waikamoi, Haiku Uka, Kula pipeline road. Drosophila truncipenna, male lekking on Cyanea with rotting flowers.

Drosophila truncipenna Waikamoi 7043
Waikamoi, Haiku Uka, Kula pipeline road. Drosophila truncipenna, courtship...
Waikamoi, Haiku Uka, Kula pipeline road. Drosophila truncipenna, courtship dance on Cyanea with rotting flowers.

Drosophila truncipenna Waikamoi 7044
Waikamoi, Haiku Uka, Kula pipeline road. Drosophila truncipenna, courtship...
Waikamoi, Haiku Uka, Kula pipeline road. Drosophila truncipenna, courtship dance on Cyanea with rotting flowers.

Waikamoi, Haiku Uka, Kula pipeline road. Drosophila truncipenna, courtship...
Waikamoi, Haiku Uka, Kula pipeline road. Drosophila truncipenna, courtship dance on Cyanea with rotting flowers.

Waikamoi, Haiku Uka, Kula pipeline road. Drosophila truncipenna.

Waikamoi, Haiku Uka, Kula pipeline road. Drosophila truncipenna, two males...
Waikamoi, Haiku Uka, Kula pipeline road. Drosophila truncipenna, two males competing for lek positions.

Lispocephala sp Waikamoi 7072
Waikamoi, Haiku Uka, Kula pipeline road. Lispocephala sp.

Lispocephala sp Waikamoi 7076
Waikamoi, Haiku Uka, Kula pipeline road. Lispocephala sp.

Lispocephala sp Waikamoi 7078
Waikamoi, Haiku Uka, Kula pipeline road. Lispocephala sp.

Waikamoi, Haiku Uka. Heed Trail. Drosophila adiastola.

Waikamoi, Haiku Uka. Heed Trail. Drosophila fasciculisetae.

Drosophila melanocephala Waikamoi 6926
Waikamoi, Haiku Uka. Heed Trail. Drosophila melanocephala.

Drosophila melanocephala Waikamoi 6929
Waikamoi, Haiku Uka. Heed Trail. Drosophila melanocephala.

Drosophila melanocephala Waikamoi 6937
Waikamoi, Haiku Uka. Heed Trail. Drosophila melanocephala.

Drosophila planitibia Waikamoi 6940
Waikamoi, Haiku Uka. Heed Trail. Drosophila planitibia.

Drosophila planitibia Waikamoi 6942
Waikamoi, Haiku Uka. Heed Trail. Drosophila planitibia.

Drosophila planitibia Waikamoi 6943
Waikamoi, Haiku Uka. Heed Trail. Drosophila planitibia.

Campsicnemus Heed Trail 1820
July 31, 2007. East Maui, Haiku Uka, Heed Trail. Campsicnemus sp.

Campsicnemus Heed Trail 1821
July 31, 2007. East Maui, Haiku Uka, Heed Trail. Campsicnemus feeding on a...
July 31, 2007. East Maui, Haiku Uka, Heed Trail. Campsicnemus feeding on a springtail.

Drosophila paucitarsus Heed Trail 1822
July 31, 2007. East Maui, Haiku Uka, Heed Trail. Drosophila lekking on...
July 31, 2007. East Maui, Haiku Uka, Heed Trail. Drosophila lekking on Melicope leaf.

Drosophila paucitarsus Heed Trail 1823
July 31, 2007. East Maui, Haiku Uka, Heed Trail. Drosophila lekking on...
July 31, 2007. East Maui, Haiku Uka, Heed Trail. Drosophila lekking on Melicope leaf.

Drosophila paucitarsus Heed Trail 1824
July 31, 2007. East Maui, Haiku Uka, Heed Trail. Drosophila lekking on...
July 31, 2007. East Maui, Haiku Uka, Heed Trail. Drosophila lekking on Melicope leaf.

Scaptomyza nr cyrtandrae Heed Trail 1833
July 31, 2007. East Maui, Haiku Uka, Heed Trail. Scaptomyza (Elmomyza) n....
July 31, 2007. East Maui, Haiku Uka, Heed Trail. Scaptomyza (Elmomyza) n. sp. nr. cyrtandrae in flower of Cyrtandra platyphylla.

Scaptomyza on Cyrtandra Waikamoi 1164
August 19, 2009. Waikamoi (Haiku Uka), Carson Trail. Scaptomyza (Elmomyza)...
August 19, 2009. Waikamoi (Haiku Uka), Carson Trail. Scaptomyza (Elmomyza) sp. feeding on a dead psyllid on Cyrtandra platyphylla.

Scaptomyza on Cyrtandra Waikamoi 1165
August 19, 2009. Waikamoi (Haiku Uka), Carson Trail. Scaptomyza (Elmomyza)...
August 19, 2009. Waikamoi (Haiku Uka), Carson Trail. Scaptomyza (Elmomyza) sp. feeding on a dead psyllid on Cyrtandra platyphylla.

Scaptomyza on Cyrtandra Waikamoi 1169
August 19, 2009. Waikamoi (Haiku Uka), Carson Trail. Scaptomyza (Elmomyza)...
August 19, 2009. Waikamoi (Haiku Uka), Carson Trail. Scaptomyza (Elmomyza) sp. feeding on a dead psyllid on Cyrtandra platyphylla.

Waikamoi, Haiku Uka, Kula Pipeline road. Drosophila clavisetae on rotting...
Waikamoi, Haiku Uka, Kula Pipeline road. Drosophila clavisetae on rotting stem of Cyanea mceldowneyi.

Campsicnemus capitulatus Makawao 1860
July 31, 2007. East Maui, Makawao FR, trail to lower Waikamoi Preserve....
July 31, 2007. East Maui, Makawao FR, trail to lower Waikamoi Preserve. Campsicnemus capitulatus on Clermontia leaf litter.

Campsicnemus capitulatus Makawao 1861
July 31, 2007. East Maui, Makawao FR, trail to lower Waikamoi Preserve....
July 31, 2007. East Maui, Makawao FR, trail to lower Waikamoi Preserve. Campsicnemus capitulatus on Clermontia leaf litter.

Campsicnemus capitulatus Makawao 1862
July 31, 2007. East Maui, Makawao FR, trail to lower Waikamoi Preserve....
July 31, 2007. East Maui, Makawao FR, trail to lower Waikamoi Preserve. Campsicnemus capitulatus on Clermontia leaf litter.

Lispocephala with prey Makawao 1863
July 31, 2007. East Maui, Makawao FR, trail to lower Waikamoi Preserve. Lispocephala flaccida or longipes with prey (Drosophila or Campsicnemus).

Lispocephala with prey Makawao 1864
July 31, 2007. East Maui, Makawao FR, trail to lower Waikamoi Preserve. Lispocephala flaccida or longipes with prey (Drosophila or Campsicnemus).

Lispocephala with prey Makawao 1865
July 31, 2007. East Maui, Makawao FR, trail to lower Waikamoi Preserve. Lispocephala flaccida or longipes with prey (Drosophila or Campsicnemus).

Hylaeus psammobius Pauwela 1151
Eleilei Point, Pauwela. Hylaeus psammobius.

Drosophila paenihamifera Hanaula 1499
Hanaula, West Maui FR. Drosophila paenihamifera, female.

Drosophila paenihamifera Hanaula 1501
Hanaula, West Maui FR. Drosophila paenihamifera, female.

Drosophila paenihamifera Hanaula 1502
Hanaula, West Maui FR. Drosophila paenihamifera, male.

Hanaula, West Maui FR. Drosophila paenihamifera, male in characteristic...
Hanaula, West Maui FR. Drosophila paenihamifera, male in characteristic front-up stance.

Mecyclothorax sp Hanaula 1494
Hanaula, West Maui FR. Mecyclothorax sp. These beetles occasionally come to...
Hanaula, West Maui FR. Mecyclothorax sp. These beetles occasionally come to Drosophila baits and I always assumed it was to prey on other insects attracted to it, but here there were several that were clearly feeding on the banana bait itself.

Scaptomyza nasalis Puu Kukui 1128
August 18, 2009. Puu Kukui Trail, 3650'. Scaptomyza (Engiscaptomyza)...
August 18, 2009. Puu Kukui Trail, 3650'. Scaptomyza (Engiscaptomyza) nasalis on Dubautia flower.

Scaptomyza nasalis Puu Kukui 1126
August 18, 2009. Puu Kukui Trail, 3650'. Scaptomyza (Engiscaptomyza)...
August 18, 2009. Puu Kukui Trail, 3650'. Scaptomyza (Engiscaptomyza) nasalis on Dubautia flower.

Scaptomyza nasalis Puu Kukui 1123
August 18, 2009. Puu Kukui Trail, 3650'. Scaptomyza (Engiscaptomyza)...
August 18, 2009. Puu Kukui Trail, 3650'. Scaptomyza (Engiscaptomyza) nasalis on Dubautia flower.

Nesophrosyne broussaisiai Puu Kukui 9424
Nesophrosyne broussaisiai, a native leafhopper that feeds on Broussaisia...
Nesophrosyne broussaisiai, a native leafhopper that feeds on Broussaisia arguta (kanawao). Puu Kukui Trail.

Megalagrion nigrohamatum Puu Kukui 2123
August 7, 2007. Puu Kukui Preserve, Kahanaiki Gulch. Megalagrion...
August 7, 2007. Puu Kukui Preserve, Kahanaiki Gulch. Megalagrion nigrohamatum nigrohamatum.

Megalagrion nigrohamatum Puu Kukui 2124
August 7, 2007. Puu Kukui Preserve, Kahanaiki Gulch. Megalagrion...
August 7, 2007. Puu Kukui Preserve, Kahanaiki Gulch. Megalagrion nigrohamatum nigrohamatum.

Megalagrion nigrohamatum Puu Kukui 2125
August 7, 2007. Puu Kukui Preserve, Kahanaiki Gulch. Megalagrion...
August 7, 2007. Puu Kukui Preserve, Kahanaiki Gulch. Megalagrion nigrohamatum nigrohamatum.

Phaeogramma hispida gall Mahinahina 4139
Phaeogramma hispida galls on Bidens sp. Mahinahina Gulch, Puu Kukui Preserve.

Hylaeus psammobius Papanalahoa 9376
Hylaeus psammobius. Papanalahoa, West Maui. This female is extruding a...
Hylaeus psammobius. Papanalahoa, West Maui. This female is extruding a droplet of nectar to concentrate it (it also contains a lot of pollen).

Hylaeus psammobius Papanalahoa 9378
Hylaeus psammobius. Papanalahoa, West Maui. This female is extruding a...
Hylaeus psammobius. Papanalahoa, West Maui. This female is extruding a droplet of nectar to concentrate it (it also contains a lot of pollen).

Hylaeus psammobius Papanalahoa 9381
Hylaeus psammobius. Papanalahoa, West Maui. This female is extruding a...
Hylaeus psammobius. Papanalahoa, West Maui. This female is extruding a droplet of nectar to concentrate it (it also contains a lot of pollen).

Tetragnatha feeding Puu Kolekole 7089
Tetragnatha sp., Puu Kolekole. This species does not build a web, as...
Tetragnatha sp., Puu Kolekole. This species does not build a web, as indicated by the spiny bristles on the legs.

Theridion grallator Puu Kolekole 7084
Puu Kolekole, Kamakou Preserve, Molokai. Theridion grallator, happyface spider.

Drosophila grimshawi Huewai 6811
Kawela, Huewai Gulch (east of Puu Kolekole road). Drosophila grimshawi.

Drosophila grimshawi Huewai 6812
Kawela, Huewai Gulch (east of Puu Kolekole road). Drosophila grimshawi.

Drosophila grimshawi Huewai 6821
Kawela, Huewai Gulch (east of Puu Kolekole road). Drosophila grimshawi.

Drosophila sodomae Huewai 6829
Kawela, Huewai Gulch (east of Puu Kolekole road). Drosophila sodomae.

Drosophila sodomae Huewai 6832
Kawela, Huewai Gulch (east of Puu Kolekole road). Drosophila sodomae.

Drosophila neoperkinsi Pepeopae 6687
Kamakou Preserve, trail to Pelekunu lookout. Drosophila neoperkinsi.

Drosophila neoperkinsi Pepeopae 6690
Kamakou Preserve, trail to Pelekunu lookout. Drosophila neoperkinsi.

Drosophila neoperkinsi Pepeopae 6691
Kamakou Preserve, trail to Pelekunu lookout. Drosophila neoperkinsi.

Megalagrion koelense Pepeopae 6654
Kamakou Preserve, trail to Pelekunu lookout. Megalagrion koelense.

Melicope leaf mines 6646
Kamakou Preserve, near Hanalilolilo trail. Leaf mines in Melicope sp. These...
Kamakou Preserve, near Hanalilolilo trail. Leaf mines in Melicope sp. These were especially abundant here, but appeared to be always narrow despite often being very long and winding.

Drosophila bostrycha Hanalilolilo 6704
Kamakou Preserve, Hanalilolilo. Drosophila bostrycha.

Drosophila bostrycha Hanalilolilo 6705
Kamakou Preserve, Hanalilolilo. Drosophila bostrycha.

Drosophila neoperkinsi Hanalilolilo 6709
Kamakou Preserve, Hanalilolilo. Drosophila neoperkinsi.

Drosophila neoperkinsi Hanalilolilo 6711
Kamakou Preserve, Hanalilolilo. Drosophila neoperkinsi.

Drosophila neoperkinsi Hanalilolilo 6712
Kamakou Preserve, Hanalilolilo. Drosophila neoperkinsi.

Hylaeus difficilis mating Kupaia 7181
A pair of native bees (Hylaeus difficilis) mating, Kupaia ridge, Molokai.

Drosophila cilifera Mokomoko 6751
Mokomoko Gulch above Kapuna spring. Drosophila cilifera.

Drosophila cilifera Mokomoko 6752
Mokomoko Gulch above Kapuna spring. Drosophila cilifera.

Drosophila cilifera Mokomoko 6758
Mokomoko Gulch above Kapuna spring. Drosophila cilifera.

Megalagrion sp Mokomoko 6728
Mokomoko Gulch above Kapuna spring. Megalagrion sp.

Megalagrion sp Mokomoko 6729
Mokomoko Gulch above Kapuna spring. Megalagrion sp.

Megalagrion sp Mokomoko 6730
Mokomoko Gulch above Kapuna spring. Megalagrion sp.

Hylaeus anthracinus wall Sandy Beach 2750
Rock and coral wall at Sandy Beach, Oahu. Hylaeus anthracinus were nesting...
Rock and coral wall at Sandy Beach, Oahu. Hylaeus anthracinus were nesting in the crevices and holes in the coral here, and apparently nowhere else along the beach; they were only found within about 100 feet of the wall. Note that many of these are not actually white coral, but black or gray rocks painted white; not sure what the point of that is.

Drosophila inedita Pia 2308
September 26, 2009. Pia Valley. Drosophila inedita.

Drosophila inedita Pia 2310
September 26, 2009. Pia Valley. Drosophila inedita.

ohia bud caterpillar 2615
Bud-feeding caterpillar on ohia. Wiliwilinui Trail.

ohia frass tube caterpillar 2613
Frass-tube caterpillar on ohia leaf. Wiliwilinui Trail.

salticid Wiliwilinui 1240
May 27, 2007. Wiliwilinui Trail, Oahu. Mike Richardson caught a number of...
May 27, 2007. Wiliwilinui Trail, Oahu. Mike Richardson caught a number of these big jumping spiders in Freycinetia axils, from the summit about halfway down the trail.

salticid Wiliwilinui 1241
May 27, 2007. Wiliwilinui Trail, Oahu. Mike Richardson caught a number of...
May 27, 2007. Wiliwilinui Trail, Oahu. Mike Richardson caught a number of these big jumping spiders in Freycinetia axils, from the summit about halfway down the trail.

Prognathogryllus sp Kaau 1714
Prognathogryllus sp. tree cricket, Kaau Crater. These are usually nocturnal;...
Prognathogryllus sp. tree cricket, Kaau Crater. These are usually nocturnal; this one was coming out at the end of the day.

Prognathogryllus sp Kaau 1715
Prognathogryllus sp. tree cricket, Kaau Crater. These are usually nocturnal;...
Prognathogryllus sp. tree cricket, Kaau Crater. These are usually nocturnal; this one was coming out at the end of the day.

Trioza sp glab Kaau 1709
Galls of Trioza sp. psyllids on Metrosideros polymorpha var. glaberrima, ohia...
Galls of Trioza sp. psyllids on Metrosideros polymorpha var. glaberrima, ohia lehua. Kaau Crater.

Trioza sp glab Kaau 1710
Galls of Trioza sp. psyllids on Metrosideros polymorpha var. glaberrima, ohia...
Galls of Trioza sp. psyllids on Metrosideros polymorpha var. glaberrima, ohia lehua. Kaau Crater.

Trioza sp glab Kaau 1711
Galls of Trioza sp. psyllids on Metrosideros polymorpha var. glaberrima, ohia...
Galls of Trioza sp. psyllids on Metrosideros polymorpha var. glaberrima, ohia lehua. Kaau Crater.

Kaau Crater 1702
Kaau Crater.

Kaau Crater 1703
Kaau Crater.

Kaau Crater pano 1702
Kaau Crater.

Trioza sp macropus Kaau 1698
Galls of Trioza sp. psyllids on Metrosideros macropus, lehua mamo. Kaau Crater.

Trioza sp macropus Kaau 1699
Galls of Trioza sp. psyllids on Metrosideros macropus, lehua mamo. Kaau Crater.

Trioza sp macropus Kaau 1700
Galls of Trioza sp. psyllids on Metrosideros macropus, lehua mamo. Kaau Crater.

Drosophila Freycinetia larva Manoa Cliff 7320
Larva of a native Drosophila living in Freycinetia (ieie) leaf axils. Manoa...
Larva of a native Drosophila living in Freycinetia (ieie) leaf axils. Manoa Cliff Trail.

Drosophila Freycinetia larva Manoa Cliff 7322
Larva of a native Drosophila living in Freycinetia (ieie) leaf axils. Manoa...
Larva of a native Drosophila living in Freycinetia (ieie) leaf axils. Manoa Cliff Trail.

Drosophila Freycinetia larva Manoa Cliff 7351
Larva of a native Drosophila living in Freycinetia (ieie) leaf axils. Manoa...
Larva of a native Drosophila living in Freycinetia (ieie) leaf axils. Manoa Cliff Trail.

Drosophila crucigera Nuuanu 0617
August 2, 2009. Above Nuuanu Pali overlook, at Charpentiera patch....
August 2, 2009. Above Nuuanu Pali overlook, at Charpentiera patch. Drosophila crucigera.

Drosophila crucigera Nuuanu 0622
August 2, 2009. Above Nuuanu Pali overlook, at Charpentiera patch....
August 2, 2009. Above Nuuanu Pali overlook, at Charpentiera patch. Drosophila crucigera.

Drosophila inedita Nuuanu 0639
August 2, 2009. Above Nuuanu Pali overlook, at Charpentiera patch....
August 2, 2009. Above Nuuanu Pali overlook, at Charpentiera patch. Drosophila inedita.

Drosophila inedita Nuuanu 0640
August 2, 2009. Above Nuuanu Pali overlook, at Charpentiera patch. Drosophila inedita.

Drosophila inedita Nuuanu 0644
August 2, 2009. Above Nuuanu Pali overlook, at Charpentiera patch. Drosophila inedita.

Drosophila moli Nuuanu 0625
August 2, 2009. Above Nuuanu Pali overlook, at Charpentiera patch. Drosophila moli (formerly included in D. lanaiensis).

Drosophila moli Nuuanu 0625a
August 2, 2009. Above Nuuanu Pali overlook, at Charpentiera patch. Drosophila moli (formerly included in D. lanaiensis).

Drosophila moli Nuuanu 0625b
August 2, 2009. Above Nuuanu Pali overlook, at Charpentiera patch. Drosophila moli (formerly included in D. lanaiensis).

Drosophila moli Nuuanu 0626

Drosophila moli Nuuanu 0628

Drosophila on sponge Nuuanu 0654
August 2, 2009. Above Nuuanu Pali overlook. This shows the density of alien...
August 2, 2009. Above Nuuanu Pali overlook. This shows the density of alien Drosophila at this site.

Drosophila punalua Nuuanu 0610
August 2, 2009. Above Nuuanu Pali overlook, at Charpentiera patch. Drosophila punalua.

Drosophila punalua Nuuanu 0612
August 2, 2009. Above Nuuanu Pali overlook, at Charpentiera patch. Drosophila punalua.

Drosophila punalua Nuuanu 0613
August 2, 2009. Above Nuuanu Pali overlook, at Charpentiera patch. Drosophila punalua.

August 2, 2009. Above Nuuanu Pali overlook, at Charpentiera patch. Drosophila punalua, on rotting Charpentiera log.

Drosophila moli, a rare species that breeds in decaying bark of Charpentiera...
Drosophila moli, a rare species that breeds in decaying bark of Charpentiera (papala). Above Nuuanu Pali lookout.

Drosophila moli Nuuanu 7250a
Drosophila moli, a rare species that breeds in decaying bark of Charpentiera...
Drosophila moli, a rare species that breeds in decaying bark of Charpentiera (papala). Above Nuuanu Pali lookout.

Drosophila moli Nuuanu eggs 7230
Eggs of Drosophila moli inserted into decaying bark of Charpentiera (papala)....
Eggs of Drosophila moli inserted into decaying bark of Charpentiera (papala). Above Nuuanu Pali lookout.

Nabis Nuuanu 0666a
August 2, 2009. Above Nuuanu Pali overlook. This nabid caught an alien...
August 2, 2009. Above Nuuanu Pali overlook. This nabid caught an alien Drosophila on a bait sponge and carried it around for some time.

Drosophila Freycinetia larva Nuuanu 7203
Larva of a native Drosophila living in Freycinetia (ieie) leaf axils. Above...
Larva of a native Drosophila living in Freycinetia (ieie) leaf axils. Above Nuuanu Pali lookout.

Drosophila Freycinetia larva Nuuanu 7204
Larva of a native Drosophila living in Freycinetia (ieie) leaf axils. Above...
Larva of a native Drosophila living in Freycinetia (ieie) leaf axils. Above Nuuanu Pali lookout.

Drosophila Freycinetia larva Nuuanu 7206
Larva of a native Drosophila living in Freycinetia (ieie) leaf axils. Above...
Larva of a native Drosophila living in Freycinetia (ieie) leaf axils. Above Nuuanu Pali lookout.

salticid Nuuanu 1798
Jumping spider, Salticidae. Mesic forest above Nuuanu Lookout, Oahu.

Megaprosternum longiceps 0544

Theridion grallator Moanalua 4058
Theridion grallator, happyface spider. Moanalua Valley.

Theridion grallator Moanalua 4063
Theridion grallator, happyface spider. Moanalua Valley.

Megaprosternum longiceps 0543

Proterhinus sp Moanalua 6252
Proterhinus sp. Kulanaahane Trail, Moanalua Valley.

Hylaeus mamo Tripler 3360
Hylaeus n. sp., Tripler ridge. No other Oahu species have the mandibles...
Hylaeus n. sp., Tripler ridge. No other Oahu species have the mandibles yellow like this.

Hylaeus mamo Tripler 3360a
Hylaeus n. sp., Tripler ridge. No other Oahu species have the mandibles...
Hylaeus n. sp., Tripler ridge. No other Oahu species have the mandibles yellow like this.

Hylaeus mamo Tripler 3364
Hylaeus n. sp., Tripler ridge. No other Oahu species have the mandibles...
Hylaeus n. sp., Tripler ridge. No other Oahu species have the mandibles yellow like this.

Hyposmocoma kahamanoa Moanalua 6245
Hyposmocoma kahamanoa, a native case-making caterpillar that lives in...
Hyposmocoma kahamanoa, a native case-making caterpillar that lives in streams. Kulanaahane Trail, Moanalua Valley.

Hyposmocoma kahamanoa Moanalua 6246
Hyposmocoma kahamanoa, a native case-making caterpillar that lives in...
Hyposmocoma kahamanoa, a native case-making caterpillar that lives in streams. Kulanaahane Trail, Moanalua Valley.

Sierola armata Moanalua 6248
Sierola armata, a native parasitic wasp, attacking a caterpillar of...
Sierola armata, a native parasitic wasp, attacking a caterpillar of Hyposmocoma kahamanoa. Kulanaahane Trail, Moanalua Valley.

Paratachardina pseudolobata Moanalua 6242
Paratachardina pseudolobata, lobate lac scale, on the native tree Myrsine...
Paratachardina pseudolobata, lobate lac scale, on the native tree Myrsine lessertiana. Kulanaahane Trail, Moanalua Valley.

Paratachardina pseudolobata Moanalua 3223
Paratachardina pseudolobata, lobate lac scale. On twigs of cultivated white...
Paratachardina pseudolobata, lobate lac scale. On twigs of cultivated white Hibiscus, Moanalua Gardens. A generalist plant pest new to Hawaii, already starting to invade native forest. When crushed, they produce a blood-red juice like cochineal. The small red dots are the immature crawler stage.

Paratachardina pseudolobata Moanalua 3224
Paratachardina pseudolobata, lobate lac scale. On twigs of cultivated white...
Paratachardina pseudolobata, lobate lac scale. On twigs of cultivated white Hibiscus, Moanalua Gardens. A generalist plant pest new to Hawaii, already starting to invade native forest. When crushed, they produce a blood-red juice like cochineal. The small red dots are the immature crawler stage.

Paratachardina pseudolobata, lobate lac scale. On twigs of cultivated white...
Paratachardina pseudolobata, lobate lac scale. On twigs of cultivated white Hibiscus, Moanalua Gardens. A generalist plant pest new to Hawaii, already starting to invade native forest. When crushed, they produce a blood-red juice like cochineal. The small red dots are the immature crawler stage.

Tripler pond 1 5817
The old mitigation ponds at Tripler, above the hose-fed "stream" where...
The old mitigation ponds at Tripler, above the hose-fed "stream" where Megalagrion xanthomelas lives as of 2014. The concrete ponds, the fence surrounding them, and even the shadecloth covering them are largely intact, though they have grass growing inside. The wooden scaffold over pond 1 (the most mauka) has partially collapsed, while the shadecloth on pond 2 has a heavy load of fallen leaves. The pump looks mostly intact from a distance, aside from one cut pipe, though due to the locked gate I couldn't check the wiring.

Tripler pond 2 5816
The old mitigation ponds at Tripler, above the hose-fed "stream" where...
The old mitigation ponds at Tripler, above the hose-fed "stream" where Megalagrion xanthomelas lives as of 2014. The concrete ponds, the fence surrounding them, and even the shadecloth covering them are largely intact, though they have grass growing inside. The wooden scaffold over pond 1 (the most mauka) has partially collapsed, while the shadecloth on pond 2 has a heavy load of fallen leaves. The pump looks mostly intact from a distance, aside from one cut pipe, though due to the locked gate I couldn't check the wiring.

Tripler pond 4 5815
The old mitigation ponds at Tripler, above the hose-fed "stream" where...
The old mitigation ponds at Tripler, above the hose-fed "stream" where Megalagrion xanthomelas lives as of 2014. The concrete ponds, the fence surrounding them, and even the shadecloth covering them are largely intact, though they have grass growing inside. The wooden scaffold over pond 1 (the most mauka) has partially collapsed, while the shadecloth on pond 2 has a heavy load of fallen leaves. The pump looks mostly intact from a distance, aside from one cut pipe, though due to the locked gate I couldn't check the wiring.

Tripler ponds pump 5818

Megalagrion xanthomelas Tripler 3933
Megalagrion xanthomelas, Tripler Army Medical Center. This is the only...
Megalagrion xanthomelas, Tripler Army Medical Center. This is the only population remaining on Oahu, and only persists in an artificial stream.

Megalagrion xanthomelas Tripler 3935
Megalagrion xanthomelas, Tripler Army Medical Center. This is the only...
Megalagrion xanthomelas, Tripler Army Medical Center. This is the only population remaining on Oahu, and only persists in an artificial stream.

Megalagrion xanthomelas Tripler 3937
Megalagrion xanthomelas, Tripler Army Medical Center. This is the only...
Megalagrion xanthomelas, Tripler Army Medical Center. This is the only population remaining on Oahu, and only persists in an artificial stream.

Tripler stream near ponds 5819
The steeper portion of the stream immediately adjacent to the old mitigation...
The steeper portion of the stream immediately adjacent to the old mitigation ponds is channelized with stones, making it easy to follow. However, a short distance down, the paving has broken up and sediment has built up on top, and the streambed is less well defined.

Tripler stream below ponds 5821
The steeper portion of the stream immediately adjacent to the old mitigation...
The steeper portion of the stream immediately adjacent to the old mitigation ponds is channelized with stones, making it easy to follow. However, a short distance down, the paving has broken up and sediment has built up on top, and the streambed is less well defined.

CRB gathering Hickam 5072
The crew of ag, forestry, and conservation people gathering to take a look at...
The crew of ag, forestry, and conservation people gathering to take a look at the coconut rhinoceros beetle infestation at Mamala Bay golf course, Joint Base Pearl Harbor-Hickam.

CRB mulch pile Hickam 5080
The main "mulch pile" where coconut rhinoceros beetles were found breeding,...
The main "mulch pile" where coconut rhinoceros beetles were found breeding, Mamala Bay golf course, Joint Base Pearl Harbor-Hickam. The name is a bit misleading, as the accumulated compost is about 200 yards long and up to 4 feet deep on the right side of the picture, where the ground slopes down to a stream - quite a lot of material. There are also some smaller piles scattered around the golf course, mainly of grass clippings.

CRB Oryctes rhinoceros Hickam 5074
Adult coconut rhinoceros beetle, Oryctes rhinoceros. Mamala Bay golf course,...
Adult coconut rhinoceros beetle, Oryctes rhinoceros. Mamala Bay golf course, Joint Base Pearl Harbor-Hickam. At the time this was taken a mesh covering had been put over the mulch pile that they're breeding in, in order to minimize the number of emerging adults that spread around while they could figure out what to do about the pile itself.

CRB Oryctes rhinoceros Hickam 5076
Larval coconut rhinoceros beetles, Oryctes rhinoceros. Mamala Bay golf...
Larval coconut rhinoceros beetles, Oryctes rhinoceros. Mamala Bay golf course, Joint Base Pearl Harbor-Hickam. These were dug out of the mulch pile without much trouble, indicating how many are present in the whole thing. The ones with smaller head capsules that are keeping on their backs rather than on their sides are a different large scarab, a flower beetle.

CRB Oryctes rhinoceros Hickam 5077
Larval coconut rhinoceros beetles, Oryctes rhinoceros. Mamala Bay golf...
Larval coconut rhinoceros beetles, Oryctes rhinoceros. Mamala Bay golf course, Joint Base Pearl Harbor-Hickam. These were dug out of the mulch pile without much trouble, indicating how many are present in the whole thing. The ones with smaller head capsules that are keeping on their backs rather than on their sides are a different large scarab, a flower beetle.

CRB trap Hickam 5073
Trap for coconut rhinoceros beetle, Oryctes rhinoceros. Mamala Bay golf...
Trap for coconut rhinoceros beetle, Oryctes rhinoceros. Mamala Bay golf course, Joint Base Pearl Harbor-Hickam.

CRB coconut damage Hickam 5081
The characteristic leaf damage caused by coconut rhinoceros beetles, Oryctes...
The characteristic leaf damage caused by coconut rhinoceros beetles, Oryctes rhinoceros. Mamala Bay golf course, Joint Base Pearl Harbor-Hickam. Palm fronds show triangular cutouts or slightly rounded cut-off tops, the result of adult beetles boring into the growing tips to feed on sap. However, since it takes months for the frond to grow out and emerge, the beetles are long gone by the time this kind of damage is visible.

CRB coconut damage Hickam 5082
The characteristic leaf damage caused by coconut rhinoceros beetles, Oryctes...
The characteristic leaf damage caused by coconut rhinoceros beetles, Oryctes rhinoceros. Mamala Bay golf course, Joint Base Pearl Harbor-Hickam. Palm fronds show triangular cutouts or slightly rounded cut-off tops, the result of adult beetles boring into the growing tips to feed on sap. However, since it takes months for the frond to grow out and emerge, the beetles are long gone by the time this kind of damage is visible.

CRB coconut damage Hickam 5085
Boreholes in coconut rachises caused by coconut rhinoceros beetles, Oryctes...
Boreholes in coconut rachises caused by coconut rhinoceros beetles, Oryctes rhinoceros. Mamala Bay golf course, Joint Base Pearl Harbor-Hickam. The adult beetles bore through the outer fronds to get to the growing shoot, where they feed on sap. This is the definitive sign of CRB infestation, and is also visible much earlier than leaf damage (though less conspicuous). This tree shows signs of at least three attacks; repeated attacks will eventually kill the tree.

Boreholes in the top of a coconut tree caused by coconut rhinoceros beetles,...
Boreholes in the top of a coconut tree caused by coconut rhinoceros beetles, Oryctes rhinoceros. Mamala Bay golf course, Joint Base Pearl Harbor-Hickam. The adult beetles bore through the outer fronds to get to the growing shoot, where they feed on sap. This is the definitive sign of CRB infestation, and is also visible much earlier than leaf damage (though less conspicuous).

Boreholes in the growing shoot of a coconut tree from coconut rhinoceros...
Boreholes in the growing shoot of a coconut tree from coconut rhinoceros beetles, Oryctes rhinoceros. Mamala Bay golf course, Joint Base Pearl Harbor-Hickam. The adult beetles bore through the outer fronds to get to the growing shoot, where they feed on sap. This is the definitive sign of CRB infestation, and is also visible much earlier than leaf damage (though less conspicuous).

Boreholes in the growing shoot of a coconut tree from coconut rhinoceros...
Boreholes in the growing shoot of a coconut tree from coconut rhinoceros beetles, Oryctes rhinoceros. Mamala Bay golf course, Joint Base Pearl Harbor-Hickam. The adult beetles bore through the outer fronds to get to the growing shoot, where they feed on sap. These beetles were caught in the act, and the picture shows how a single bore causes holes in multiple rachises.

Boreholes in the growing shoot of a coconut tree from coconut rhinoceros...
Boreholes in the growing shoot of a coconut tree from coconut rhinoceros beetles, Oryctes rhinoceros. Mamala Bay golf course, Joint Base Pearl Harbor-Hickam. The adult beetles bore through the outer fronds to get to the growing shoot, where they feed on sap. This shows how a single bore causes holes in multiple rachises.

Boreholes in the growing shoot of a coconut tree from coconut rhinoceros beetles, Oryctes rhinoceros. Mamala Bay golf course, Joint Base Pearl Harbor-Hickam. The adult beetles bore through the outer fronds to get to the growing shoot, where they feed on sap. This shows how a single bore causes holes in multiple rachises.

Tetreuaresta obscuriventris Aiea 6107
Tetreuaresta obscuriventris, a tephritid fruit fly introduced as a biocontrol...
Tetreuaresta obscuriventris, a tephritid fruit fly introduced as a biocontrol for Elephantopus mollis (the flower it's sitting on). Elephantopus has completely taken over the Aiea Ridge Trail where it wasn't present before, so hopefully the fly will be able to bring it under control.

Tetreuaresta obscuriventris Aiea 6108
Tetreuaresta obscuriventris, a tephritid fruit fly introduced as a biocontrol...
Tetreuaresta obscuriventris, a tephritid fruit fly introduced as a biocontrol for Elephantopus mollis (the flower it's sitting on). Elephantopus has completely taken over the Aiea Ridge Trail where it wasn't present before, so hopefully the fly will be able to bring it under control.

Vanessa tameamea Waihee 5885
Vanessa tameamea, Kamehameha butterfly. Waihee Valley.

Vanessa tameamea Waihee 5886
Vanessa tameamea, Kamehameha butterfly. Waihee Valley.

Vanessa tameamea Waihee 5887
Vanessa tameamea, Kamehameha butterfly. Waihee Valley.

Tetragnatha sp Manana 2194
Tetragnatha sp., Manana Trail.

Tetragnatha sp Manana 2195
Tetragnatha sp., Manana Trail.

Teleonemia scrupulosa Kalaeloa 8988
Teleonemia scrupulosa, lantana lace bug. Pearl Harbor National Wildlife...
Teleonemia scrupulosa, lantana lace bug. Pearl Harbor National Wildlife Refuge, Kalaeloa section. Introduced as a biocontrol for lantana, this bug surprisingly attacks Myoporum stellatum at Barbers Point, yet doesn't feed on M. sandwicense. It's been known to do so since at least the 1960's, but is currently undergoing an outbreak and causing noticeable damage.

Teleonemia scrupulosa Kalaeloa 8991
Teleonemia scrupulosa, lantana lace bug. Pearl Harbor National Wildlife...
Teleonemia scrupulosa, lantana lace bug. Pearl Harbor National Wildlife Refuge, Kalaeloa section. Introduced as a biocontrol for lantana, this bug surprisingly attacks Myoporum stellatum at Barbers Point, yet doesn't feed on M. sandwicense. It's been known to do so since at least the 1960's, but is currently undergoing an outbreak and causing noticeable damage.

Teleonemia scrupulosa Kalaeloa 8992
Teleonemia scrupulosa, lantana lace bug. Pearl Harbor National Wildlife...
Teleonemia scrupulosa, lantana lace bug. Pearl Harbor National Wildlife Refuge, Kalaeloa section. Introduced as a biocontrol for lantana, this bug surprisingly attacks Myoporum stellatum at Barbers Point, yet doesn't feed on M. sandwicense. It's been known to do so since at least the 1960's, but is currently undergoing an outbreak and causing noticeable damage.

Megalagrion hawaiiensis Poamoho 3109
Megalagrion hawaiiensis, pinao ula. Koolau Summit Trail near Poamoho.

Megalagrion hawaiiensis Poamoho 3112
Megalagrion hawaiiensis, pinao ula. Koolau Summit Trail near Poamoho.

Megalagrion koelense Poamoho 3078
Megalagrion koelense, pinao ula. Poamoho Trail.

Colobopyga pritchardiae Poamoho 3082
Colobopyga pritchardiae, loulu scale. Poamoho Trail. These native scales...
Colobopyga pritchardiae, loulu scale. Poamoho Trail. These native scales are found only on the stems and bracts of the inflorescence of the native Pritchardia palms.

Colobopyga pritchardiae Poamoho 3083
Colobopyga pritchardiae, loulu scale. Poamoho Trail. These native scales...
Colobopyga pritchardiae, loulu scale. Poamoho Trail. These native scales are found only on the stems and bracts of the inflorescence of the native Pritchardia palms.

Trupanea gall Poamoho 2423
Trupanea gall with old pupa in Dubautia laxa, Poamoho Trail.

Megalagrion oahuense Schofield 2480
Megalagrion oahuense, pinao ula. Schofield-Waikane Trail. This species has...
Megalagrion oahuense, pinao ula. Schofield-Waikane Trail. This species has completely terrestrial nymphs, which live in wet mats of uluhe.

Megalagrion oahuense Schofield 2482
Megalagrion oahuense, pinao ula. Schofield-Waikane Trail. This species has...
Megalagrion oahuense, pinao ula. Schofield-Waikane Trail. This species has completely terrestrial nymphs, which live in wet mats of uluhe.

Drosophila oahuensis Opaeula 6286
Drosophila oahuensis, Lower Opaeula.

Drosophila substenoptera Opaeula 1930
Drosophila substenoptera. Opaeula, northern Koolau range.

Drosophila substenoptera Opaeula 6285
Drosophila substenoptera, Lower Opaeula.

Drosophila turbata Opaeula 6283
Drosophila turbata, Lower Opaeula.

Prognathogryllus sp Opaeula 6270
Prognathogryllus sp., Lower Opaeula. These tree crickets can often be heard...
Prognathogryllus sp., Lower Opaeula. These tree crickets can often be heard but they are rarely seen.

Prognathogryllus sp Opaeula 6275
Prognathogryllus sp., Lower Opaeula. These tree crickets can often be heard...
Prognathogryllus sp., Lower Opaeula. These tree crickets can often be heard but they are rarely seen.

Prognathogryllus sp Opaeula 6277
Prognathogryllus sp., Lower Opaeula. These tree crickets can often be heard...
Prognathogryllus sp., Lower Opaeula. These tree crickets can often be heard but they are rarely seen.

Drosophila oahuensis Koloa 4892
Drosophila oahuensis, Koloa.

Vanessa tameamea Opaeula 6303
Vanessa tameamea, Kamehameha butterfly, feeding on mamaki (Pipturus albidus)....
Vanessa tameamea, Kamehameha butterfly, feeding on mamaki (Pipturus albidus). Lower Opaeula.

Drosophila imitator Koloa 9711
Drosophila imitator. Koloa.

Drosophila n sp Koloa 3744
Koloa, northern Koolau range. Undescribed picture wing Drosophila, in the...
Koloa, northern Koolau range. Undescribed picture wing Drosophila, in the varipennis species complex. All previously-known members of this group are from Maui Nui.

Drosophila nr truncipenna Koloa 9714
Drosophila n. sp. nr. truncipenna, female. Koloa.

Drosophila nr truncipenna Koloa 9715
Drosophila n. sp. nr. truncipenna, female. Koloa.

Drosophila nr truncipenna Koloa 9720
Drosophila n. sp. nr. truncipenna, female. Koloa.

Drosophila oahuensis Koloa 3704
Drosophila oahuensis. Koloa, northern Koolau range.

Drosophila oahuensis Koloa 3705
Drosophila oahuensis. Koloa, northern Koolau range.

Drosophila sp courtship Koloa 5290
Drosophila sp. courtship, Koloa. The male is bending his abdomen sideways...
Drosophila sp. courtship, Koloa. The male is bending his abdomen sideways while holding up his front legs and circling the female.

Ceroplastes rubens Koloa 6194
Pink wax scale, Ceroplastes rubens, on ohia. Koloa, Koolau summit. These...
Pink wax scale, Ceroplastes rubens, on ohia. Koloa, Koolau summit. These seem to be unusually abundant now, though still not in large numbers.

Ceroplastes rubens Koloa 6196
Pink wax scale, Ceroplastes rubens, on ohia. Koloa, Koolau summit. These...
Pink wax scale, Ceroplastes rubens, on ohia. Koloa, Koolau summit. These seem to be unusually abundant now, though still not in large numbers.

Vanessa tameamea Koloa
Vanessa tameamea, Kamehameha butterfly. Koloa, on the Koolau Summit Trail....
Vanessa tameamea, Kamehameha butterfly. Koloa, on the Koolau Summit Trail. This area was quite far from its host plants, mamaki and opuhe. Photo by Kaleo Wong.

Drosophila n. sp. nr. truncipenna, male. Koloa.

Megalagrion oahuense Koloa 1819
Megalagrion oahuense. Koloa, northern Koolau range. This unusual species...
Megalagrion oahuense. Koloa, northern Koolau range. This unusual species breeds in damp uluhe litter instead of water, and is relatively common in the northern Koolaus.

Drosophila n. sp. nr. truncipenna, female. Koloa. The camera caught this...
Drosophila n. sp. nr. truncipenna, female. Koloa. The camera caught this fly as it seemed to be jumping to take off, but it ended up staying on the sponge.

Megalagrion sp Laie 2940
Megalagrion sp., painiu. Laie Trail.

caterpillar Palawai 3966

Drosophila flexipes Pualii 5356
Drosophila flexipes, Pualii.

Drosophila flexipes Pualii 5362
Drosophila flexipes, Pualii.

Drosophila flexipes Pualii 5364
Drosophila flexipes, Pualii.

Drosophila amb-mon courtship Pualii 5328
Drosophila ambochila male trying (unsuccessfully) to court a female D....
Drosophila ambochila male trying (unsuccessfully) to court a female D. montgomeryi, Pualii. The two species have different courtship dances so it's unlikely that they hybridize very often.

Drosophila amb-mon courtship Pualii 5352
Drosophila ambochila male trying (unsuccessfully) to court a female D....
Drosophila ambochila male trying (unsuccessfully) to court a female D. montgomeryi, Pualii. The two species have different courtship dances so it's unlikely that they hybridize very often.

Drosophila ambochila Pualii 5332
Drosophila ambochila on Urera kaalae, Pualii.

Drosophila ambochila Pualii 5355
Drosophila ambochila, Pualii.

Drosophila inedita Pualii 5333
Drosophila inedita, Pualii. An unusual mutant form lacking most of the...
Drosophila inedita, Pualii. An unusual mutant form lacking most of the apical wing mark.

Drosophila montgomeryi Pualii 5319
Drosophila montgomeryi on Urera kaalae, Pualii. Note the eggs laid in the...
Drosophila montgomeryi on Urera kaalae, Pualii. Note the eggs laid in the rotting bark, indicated by the white respiratory filaments.

Drosophila montgomeryi Pualii 5320
Drosophila montgomeryi on Urera kaalae, Pualii. Note the eggs laid in the...
Drosophila montgomeryi on Urera kaalae, Pualii. Note the eggs laid in the rotting bark, indicated by the white respiratory filaments.

Drosophila montgomeryi Pualii 5322
Drosophila montgomeryi ovipositing in a rotting trunk of Urera kaalae, Pualii.

Drosophila montgomeryi ovipositing in a rotting trunk of Urera kaalae, Pualii.

Drosophila montgomeryi on Urera kaalae, Pualii.

Drosophila montgomeryi eggs in a rotting trunk of Urera kaalae, Pualii.

Drosophila wikstroemiae Pualii 5338
Drosophila wikstroemiae, Pualii. Two males end-to-end; not sure exactly what...
Drosophila wikstroemiae, Pualii. Two males end-to-end; not sure exactly what they're doing like this.

Drosophila hexachaetae Palikea 1778
September 23, 2009. Palikea exclosure, east of the summit ridgeline....
September 23, 2009. Palikea exclosure, east of the summit ridgeline. Drosophila hexachaetae, at the Cyanea grimesiana exclosure.

Drosophila punalua Palikea 1783
September 23, 2009. Palikea exclosure, east of the summit ridgeline....
September 23, 2009. Palikea exclosure, east of the summit ridgeline. Drosophila punalua, at the Cyanea grimesiana exclosure.

Eupithecia Palikea 1918
September 23, 2009. Palikea exclosure, east of the summit ridgeline....
September 23, 2009. Palikea exclosure, east of the summit ridgeline. Eupithecia carnivorous caterpillar resting on a Charpentiera leaf in the Cyanea grimesiana exclosure.

Mapsidius charpentierii Palikea 2032
September 24, 2009. Palikea exclosure, east of the summit ridgeline. Mapsidius charpentierii, a stem-boring caterpillar that was heavily damaging Charpentiera saplings in the Cyanea grimesiana exclosure.

Mapsidius charpentierii Palikea 2034
September 24, 2009. Palikea exclosure, east of the summit ridgeline. Mapsidius charpentierii, a stem-boring caterpillar that was heavily damaging Charpentiera saplings in the Cyanea grimesiana exclosure.

Drosophila crucigera Pualii 5339
Drosophila crucigera ovipositing in a rotting stem of Pleomele forbesii, Pualii.

Theridion grallator Palikea 1860
September 23, 2009. Palikea exclosure, east of the summit ridgeline. Theridion grallator, happyface spider. These were very abundant in the area.

Theridion grallator Palikea 1865
September 23, 2009. Palikea exclosure, east of the summit ridgeline. Theridion grallator, happyface spider. These were very abundant in the area.

Theridion grallator Palikea 1866
September 23, 2009. Palikea exclosure, east of the summit ridgeline. Theridion grallator, happyface spider. These were very abundant in the area.

Charpentiera caterpillar damage Palawai 5016
Caterpillar feeding damage on Charpentiera tomentosa, papala. Palawai Gulch....
Caterpillar feeding damage on Charpentiera tomentosa, papala. Palawai Gulch. This appears to be different from the tip-feeding Mapsidius charpentierii which has severely affected this species in the same area.

Charpentiera caterpillar damage Palawai 5018
Caterpillar feeding damage on Charpentiera tomentosa, papala. Palawai Gulch....
Caterpillar feeding damage on Charpentiera tomentosa, papala. Palawai Gulch. This appears to be different from the tip-feeding Mapsidius charpentierii which has severely affected this species in the same area.

Theridion grallator Palawai 5011
Theridion grallator (happyface spider), Palawai Gulch.

Theridion grallator Palawai 5012
Theridion grallator (happyface spider), Palawai Gulch.

Theridion grallator Palawai 5013
Theridion grallator (happyface spider), Palawai Gulch.

Theridion sp Palawai 5027
Theridion sp., Palawai Gulch.

Theridion sp Palawai 5029
Theridion sp., Palawai Gulch.

Theridion sp Palawai 5030
Theridion sp., Palawai Gulch.

Drosophila substenoptera Palikea 2104
September 24, 2009. Palikea exclosure, east of the summit ridgeline. Drosophila substenoptera, at the Pisonia just below the end of the banyan trail. This individual was particularly persistent, returning within seconds of being scared off by the camera and staying for over an hour between visits to the site.

Drosophila substenoptera Palikea 2102
September 24, 2009. Palikea exclosure, east of the summit ridgeline. Drosophila substenoptera, at the Pisonia just below the end of the banyan trail. This individual was particularly persistent, returning within seconds of being scared off by the camera and staying for over an hour between visits to the site.

Drosophila substenoptera Palikea 2063
September 24, 2009. Palikea exclosure, east of the summit ridgeline. Drosophila substenoptera, at the Pisonia just below the end of the banyan trail. This individual was particularly persistent, returning within seconds of being scared off by the camera and staying for over an hour between visits to the site.

Drosophila substenoptera Palikea 2093 edit

September 23, 2009. Palikea exclosure, east of the summit ridgeline. Drosophila substenoptera, at the Pisonia just below the end of the banyan trail.

September 23, 2009. Palikea exclosure, east of the summit ridgeline. Drosophila substenoptera, at the Pisonia just below the end of the banyan trail. On the right is D. punalua.

September 23, 2009. Palikea exclosure, east of the summit ridgeline. Drosophila substenoptera, at the Pisonia just below the end of the banyan trail. On the left is D. punalua.

Scaptomyza Pisonia Palikea 4629
Scaptomyza (Elmomyza) sp., on fruit of Pisonia brunoniana. Palikea. This...
Scaptomyza (Elmomyza) sp., on fruit of Pisonia brunoniana. Palikea. This undescribed species breeds on the fruits of Pisonia spp. (papala kepau). The larvae crawl on or even through the sticky resin and feed on it, and they are apparently mostly able to avoid getting stuck as adults. Two eggs are visible on this fruit.

Scaptomyza Pisonia Palikea 4632
Scaptomyza (Elmomyza) sp., on fruit of Pisonia brunoniana. Palikea. This...
Scaptomyza (Elmomyza) sp., on fruit of Pisonia brunoniana. Palikea. This undescribed species breeds on the fruits of Pisonia spp. (papala kepau). The larvae crawl on or even through the sticky resin and feed on it, and they are apparently mostly able to avoid getting stuck as adults.

Scaptomyza Pisonia Palikea 4633
Scaptomyza (Elmomyza) sp., lekking near fruit of Pisonia brunoniana. Palikea....
Scaptomyza (Elmomyza) sp., lekking near fruit of Pisonia brunoniana. Palikea. This undescribed species breeds on the fruits of Pisonia spp. (papala kepau). The larvae crawl on or even through the sticky resin and feed on it, and they are apparently mostly able to avoid getting stuck as adults.

Scaptomyza (Elmomyza) sp., on fruit of Pisonia brunoniana. Palikea. This...
Scaptomyza (Elmomyza) sp., on fruit of Pisonia brunoniana. Palikea. This undescribed species breeds on the fruits of Pisonia spp. (papala kepau). The larvae crawl on or even through the sticky resin and feed on it, and they are apparently mostly able to avoid getting stuck as adults (though not always as evidenced by the one on the left).

Scaptomyza (Elmomyza) sp., on fruit of Pisonia brunoniana. Palikea. This undescribed species breeds on the fruits of Pisonia spp. (papala kepau). The larvae crawl on or even through the sticky resin and feed on it, and they are apparently mostly able to avoid getting stuck as adults.

Sierola sp Palikea 4630
Sierola sp., on Labordia. Palikea.

Sierola sp Palikea 4631
Sierola sp., on Labordia. Palikea.

Drosophila punalua Palikea 1722
September 22, 2009. Palikea exclosure, east of the summit ridgeline. Drosophila punalua.

Drosophila ambochila Palikea 1927
September 23, 2009. Palikea exclosure, east of the summit ridgeline. Drosophila ambochila? Originally thought to be D. montgomeryi, since it was found at the Urera sponges just below the large Ficus. However, it lacks the dark stripes on the mesonotum and anepisternum that D. montgomeryi has.

Drosophila ambochila Palikea 1928
September 23, 2009. Palikea exclosure, east of the summit ridgeline. Drosophila ambochila? Originally thought to be D. montgomeryi, since it was found at the Urera sponges just below the large Ficus. However, it lacks the dark stripes on the mesonotum and anepisternum that D. montgomeryi has.

Drosophila ambochila Palikea 1932
September 23, 2009. Palikea exclosure, east of the summit ridgeline. Drosophila ambochila? Originally thought to be D. montgomeryi, since it was found at the Urera sponges just below the large Ficus. However, it lacks the dark stripes on the mesonotum and anepisternum that D. montgomeryi has.

Drosophila gradata Palikea 1993
September 24, 2009. Palikea exclosure, east of the summit ridgeline. Drosophila gradata, at the Urera sponges just below the large Ficus.

Drosophila gradata Palikea 1994
September 24, 2009. Palikea exclosure, east of the summit ridgeline. Drosophila gradata, at the Urera sponges just below the large Ficus.

Drosophila gradata Palikea 1995
September 24, 2009. Palikea exclosure, east of the summit ridgeline. Drosophila gradata, at the Urera sponges just below the large Ficus.

Drosophila hemipeza Palikea 1940
September 23, 2009. Palikea exclosure, east of the summit ridgeline. Drosophila hemipeza, at the Urera sponges just below the large Ficus.

Drosophila hemipeza Palikea 1947
September 23, 2009. Palikea exclosure, east of the summit ridgeline. Drosophila hemipeza, at the Urera sponges just below the large Ficus.

Drosophila hemipeza Palikea 2013
September 24, 2009. Palikea exclosure, east of the summit ridgeline. Drosophila hemipeza, at the Urera sponges just below the large Ficus.

Drosophila hexachaetae Palikea 2008
September 24, 2009. Palikea exclosure, east of the summit ridgeline. Drosophila hexachaetae, at the Urera sponges just below the large Ficus.

Drosophila punalua Palikea 1922
September 23, 2009. Palikea exclosure, east of the summit ridgeline. Drosophila punalua, at the Urera sponges just below the large Ficus.

September 23, 2009. Palikea exclosure, east of the summit ridgeline. Drosophila substenoptera, at the Urera sponges just below the large Ficus.

Drosophila craddockae Palikea 4100
Drosophila craddockae, Palikea.

Drosophila gradata Kahanahaiki 3999
Drosophila gradata. Kahanahaiki.

Drosophila gradata Kahanahaiki 4002
Drosophila gradata. Kahanahaiki.

Drosophila hemipeza. Palikea. This species is particularly docile.

Drosophila predation Kahanahaiki 4007
Spider feeding on a Drosophila at a bait sponge. Kahanahaiki.

Drosophila substenoptera. Palikea.

Philodoria pipturus Palikea 4941
Philodoria pipturus, Puu Palikea. These native leaf-mining moths were quite...
Philodoria pipturus, Puu Palikea. These native leaf-mining moths were quite abundant for this brief period, and seem to be seasonal.

Philodoria pipturus Palikea 4943
Philodoria pipturus, Puu Palikea. These native leaf-mining moths were quite...
Philodoria pipturus, Puu Palikea. These native leaf-mining moths were quite abundant for this brief period, and seem to be seasonal.

Philodoria pipturus Palikea 4944
Philodoria pipturus, Puu Palikea. These native leaf-mining moths were quite...
Philodoria pipturus, Puu Palikea. These native leaf-mining moths were quite abundant for this brief period, and seem to be seasonal.

Philodoria pipturus Palikea 4944a

Drosophila suzukii Palikea 1882
September 23, 2009. Palikea exclosure, east of the summit ridgeline. Drosophila suzukii, resting on a thimbleberry fruit (maybe after ovipositing?).

Drosophila hexachaetae Palikea 2060
September 24, 2009. Palikea exclosure, east of the summit ridgeline. Drosophila hexachaetae, at the Urera sponges just below the large Ficus.

September 24, 2009. Palikea exclosure, east of the summit ridgeline. Drosophila hexachaetae, at the Urera sponges just below the large Ficus.

Rhyncogonus howarthi Palikea 1731
September 22, 2009. Palikea exclosure, east of the summit ridgeline. Rhyncogonus howarthi, at night near the camp. There was extensive feeding sign on many different plants, especially Kadua terminalis, and possibly R. fordi was also present.

Rhyncogonus howarthi Palikea 1734
September 22, 2009. Palikea exclosure, east of the summit ridgeline. Rhyncogonus howarthi, at night near the camp. There was extensive feeding sign on many different plants, especially Kadua terminalis, and possibly R. fordi was also present.

Rhyncogonus howarthi Palikea 1735
September 22, 2009. Palikea exclosure, east of the summit ridgeline. Rhyncogonus howarthi, at night near the camp. There was extensive feeding sign on many different plants, especially Kadua terminalis, and possibly R. fordi was also present.

September 22, 2009. Palikea exclosure, east of the summit ridgeline. Theridion grallator, happyface spider, at night near the camp. These were very abundant in the area.

Drosophila craddockae Malaekahana 5436
Drosophila craddockae, Malaekahana Trail.

Drosophila craddockae Malaekahana 5437
Drosophila craddockae, Malaekahana Trail.

Drosophila ambochila Ekahanui 0992
Ekahanui Gulch, Honouliuli. Drosophila ambochila.

Drosophila ambochila Ekahanui 0994
Ekahanui Gulch, Honouliuli. Drosophila ambochila.

Bidens stem borer Puu Kaua 2840
Stem borer pupa in Bidens torta, kookoolau. Puu Kaua trail, Honouliuli.

Toxorhynchites brevipalpis Kaluaa 6536
Toxorhynchites brevipalpis, a predatory, non-biting mosquito introduced in an...
Toxorhynchites brevipalpis, a predatory, non-biting mosquito introduced in an attempt to control biting mosquitos. Unfortunately they never really took off and it's fairly rare to see them. But when you do, they're quite a striking sight - they're big (the body is over half an inch long, and the span of the legs is close to an inch), metallic blue, strong fliers, and make a buzz similar to a regular mosquito but audible from at least five feet away. Two of them kept coming back to a particular paperbark tree near the start of the Kaluaa trail (a pretty dry area) for some reason.

Paurotriozana adaptata Kaluaa 1676
Galls and nymphs of Paurotriozana adaptata, an endemic psyllid found only on...
Galls and nymphs of Paurotriozana adaptata, an endemic psyllid found only on the last tree of Cryptocarya oahuensis, holio. Kaluaa Gulch. Another species of holio, C. mannii (with which this was formerly synonymized), occurs on Kauai but the psyllid does not.

Paurotriozana adaptata Kaluaa 1677
Galls and nymphs of Paurotriozana adaptata, an endemic psyllid found only on...
Galls and nymphs of Paurotriozana adaptata, an endemic psyllid found only on the last tree of Cryptocarya oahuensis, holio. Kaluaa Gulch. Another species of holio, C. mannii (with which this was formerly synonymized), occurs on Kauai but the psyllid does not.

Paurotriozana adaptata Kaluaa 1679
Galls and nymphs of Paurotriozana adaptata, an endemic psyllid found only on...
Galls and nymphs of Paurotriozana adaptata, an endemic psyllid found only on the last tree of Cryptocarya oahuensis, holio. Kaluaa Gulch. Another species of holio, C. mannii (with which this was formerly synonymized), occurs on Kauai but the psyllid does not.

Paurotriozana adaptata, an endemic psyllid that forms galls on Cryptocarya...
Paurotriozana adaptata, an endemic psyllid that forms galls on Cryptocarya oahuensis (holio). This is the last tree of that species, but the Paurotriozana still survives on it.

Drosophila ambochila Kaluaa 5259
Drosophila ambochila courtship, Central Kaluaa Gulch. Note that the dance of...
Drosophila ambochila courtship, Central Kaluaa Gulch. Note that the dance of the male is significantly different from that of the very similar-looking and sympatric D. montgomeryi, where the male bends his abdomen completely under his body.

Drosophila ambochila Kaluaa 5260
Drosophila ambochila courtship, Central Kaluaa Gulch. Note that the dance of...
Drosophila ambochila courtship, Central Kaluaa Gulch. Note that the dance of the male is significantly different from that of the very similar-looking and sympatric D. montgomeryi, where the male bends his abdomen completely under his body.

Drosophila ambochila Kaluaa 5261
Drosophila ambochila courtship, Central Kaluaa Gulch. Note that the dance of...
Drosophila ambochila courtship, Central Kaluaa Gulch. Note that the dance of the male is significantly different from that of the very similar-looking and sympatric D. montgomeryi, where the male bends his abdomen completely under his body.

Drosophila divaricata Kaluaa 5243
Drosophila divaricata, Central Kaluaa Gulch.

Xyleborus sp Kaluaa 4667
Xyleborus sp. boring in Pipturus albidus (mamaki). Central Kaluaa Gulch....
Xyleborus sp. boring in Pipturus albidus (mamaki). Central Kaluaa Gulch. These beetles (probably introduced) were associated with serious dieback of mamaki in this gulch, though whether they were the cause or were attacking dying trees wasn't clear.

Xyleborus sp Kaluaa 4668
Damage from Xyleborus sp. boring in Pipturus albidus (mamaki). Central...
Damage from Xyleborus sp. boring in Pipturus albidus (mamaki). Central Kaluaa Gulch. These beetles (probably introduced) were associated with serious dieback of mamaki in this gulch, though whether they were the cause or were attacking dying trees wasn't clear.

Drosophila ambochila, Kaluaa Gulch.

January 26, 2010. Kaluaa Gulch. Drosophila ambochila.

Drosophila distinguenda Kaluaa 4164
January 26, 2010. Kaluaa Gulch. Drosophila distinguenda, with D. ambochila...
January 26, 2010. Kaluaa Gulch. Drosophila distinguenda, with D. ambochila on the lower left.

Drosophila inedita Kaluaa 4162
January 26, 2010. Kaluaa Gulch. Drosophila inedita.

Drosophila inedita Kaluaa 4170
January 26, 2010. Kaluaa Gulch. Drosophila inedita.

Drosophila inedita Kaluaa 4173
January 26, 2010. Kaluaa Gulch. Drosophila inedita.

Drosophila spp Kaluaa 4207
January 26, 2010. Kaluaa Gulch. Drosophila inedita (left) and D. ambochila...
January 26, 2010. Kaluaa Gulch. Drosophila inedita (left) and D. ambochila (right).

Drosophila spp Kaluaa 4208
January 26, 2010. Kaluaa Gulch. Drosophila inedita (left) and D. ambochila...
January 26, 2010. Kaluaa Gulch. Drosophila inedita (left) and D. ambochila (right).

Scaptomyza Pisonia Kaluaa 4683
Scaptomyza (Elmomyza) sp., on fruit of Pisonia umbellifera. Central Kaluaa...
Scaptomyza (Elmomyza) sp., on fruit of Pisonia umbellifera. Central Kaluaa Gulch 1. This undescribed species breeds on the fruits of Pisonia spp. (papala kepau). The larvae crawl on or even through the sticky resin and feed on it, apparently mostly able to avoid getting stuck as adults.

Scaptomyza Pisonia Kaluaa 4688
Scaptomyza (Elmomyza) sp., on fruit of Pisonia umbellifera. Central Kaluaa...
Scaptomyza (Elmomyza) sp., on fruit of Pisonia umbellifera. Central Kaluaa Gulch 1. This undescribed species breeds on the fruits of Pisonia spp. (papala kepau). The larvae crawl on or even through the sticky resin and feed on it, apparently mostly able to avoid getting stuck as adults.

Scaptomyza Pisonia Kaluaa 4690
Scaptomyza (Elmomyza) sp., on fruit of Pisonia umbellifera. Central Kaluaa...
Scaptomyza (Elmomyza) sp., on fruit of Pisonia umbellifera. Central Kaluaa Gulch 1. This undescribed species breeds on the fruits of Pisonia spp. (papala kepau). The larvae crawl on or even through the sticky resin and feed on it, apparently mostly able to avoid getting stuck as adults.

Scaptomyza Pisonia Kaluaa 4709 stack

Vanessa tameamea Kaluaa 5421
Kamehameha butterfly (Vanessa tameamea) caterpillar on a leaf of Urera...
Kamehameha butterfly (Vanessa tameamea) caterpillar on a leaf of Urera kaalae. Kaluaa Gulch.

Drosophila ambochila feeding on the liquid oozing from a dying Urera glabra...
Drosophila ambochila feeding on the liquid oozing from a dying Urera glabra tree. Central Kaluaa Gulch. The tree was killed by scolytid beetles and heavily infected with fungus, making it apparently unsuitable as a Drosophila breeding site, but the adults were still feeding on the exudate.

Drosophila ambochila. Central Kaluaa Gulch, Honouliuli.

Drosophila inedita. Central Kaluaa Gulch, Honouliuli.

Drosophila montgomeryi Kaluaa 4652
Drosophila montgomeryi. Central Kaluaa Gulch.

Drosophila montgomeryi Kaluaa 4664
Drosophila montgomeryi. Central Kaluaa Gulch. This male was patrolling a...
Drosophila montgomeryi. Central Kaluaa Gulch. This male was patrolling a territory on a rotting branch of Urera kaalae, one of their host plants.

Drosophila montgomeryi Kaluaa 4669
Drosophila montgomeryi. Central Kaluaa Gulch. These two males were fighting...
Drosophila montgomeryi. Central Kaluaa Gulch. These two males were fighting for lekking territory on a rotting branch of Urera kaalae, one of their host plants.

Drosophila montgomeryi. Central Kaluaa Gulch. These two males were fighting...
Drosophila montgomeryi. Central Kaluaa Gulch. These two males were fighting for lekking territory on a rotting branch of Urera kaalae, one of their host plants.

Drosophila montgomeryi. Central Kaluaa Gulch, Honouliuli.

Stegana coleoptrata Kaluaa 4715
Stegana coleoptrata, attracted to a rotting Urera kaalae branch in Central...
Stegana coleoptrata, attracted to a rotting Urera kaalae branch in Central Kaluaa Gulch 1.

Stegana coleoptrata Kaluaa 4717
Stegana coleoptrata, attracted to a rotting Urera kaalae branch in Central...
Stegana coleoptrata, attracted to a rotting Urera kaalae branch in Central Kaluaa Gulch 1.

Stegana coleoptrata Kaluaa 4718
Stegana coleoptrata, attracted to a rotting Urera kaalae branch in Central...
Stegana coleoptrata, attracted to a rotting Urera kaalae branch in Central Kaluaa Gulch 1.

Stegana coleoptrata, an introduced drosophilid. Central Kaluaa Gulch.

Dictyophorodelphax mirabilis Hapapa 5767
Dictyophorodelphax mirabilis, trail to Puu Hapapa. The gut extends up...
Dictyophorodelphax mirabilis, trail to Puu Hapapa. The gut extends up through the horn and back down, and may play a role in detoxifying the milky sap of the akoko that it feeds on.

Dictyophorodelphax mirabilis Hapapa 5768
Dictyophorodelphax mirabilis, trail to Puu Hapapa. The gut extends up...
Dictyophorodelphax mirabilis, trail to Puu Hapapa. The gut extends up through the horn and back down, and may play a role in detoxifying the milky sap of the akoko that it feeds on.

Dictyophorodelphax mirabilis Hapapa 2600
Dictyophorodelphax mirabilis, the akoko planthopper, on Euphorbia...
Dictyophorodelphax mirabilis, the akoko planthopper, on Euphorbia (Chamaesyce) multiformis. Puu Hapapa trail, Honouliuli. These are usually fairly rare but can occasionally be very abundant.

Dictyophorodelphax mirabilis, the akoko planthopper, on Euphorbia...
Dictyophorodelphax mirabilis, the akoko planthopper, on Euphorbia (Chamaesyce) multiformis. Puu Hapapa trail, Honouliuli. These are usually fairly rare but can occasionally be very abundant.

Orthotylus sp Hapapa 2603
Orthotylus sp. nymph on Euphorbia (Chamaesyce) multiformis. Puu Hapapa...
Orthotylus sp. nymph on Euphorbia (Chamaesyce) multiformis. Puu Hapapa trail, Honouliuli.

Drosophila divaricata North Kaluaa 4604
Drosophila divaricata. North Kaluaa Gulch.

Drosophila divaricata North Kaluaa 4605
Drosophila divaricata. North Kaluaa Gulch.

Drosophila divaricata North Kaluaa 4606
Drosophila divaricata. North Kaluaa Gulch.

Drosophila divaricata Kaluaa 5196
Drosophila divaricata, North Kaluaa Gulch.

Drosophila divaricata Kaluaa 5203
Drosophila divaricata, North Kaluaa Gulch.

Drosophila divaricata, North Kaluaa Gulch.

Drosophila montgomeryi, north Kaluaa Gulch. Second from left is D. ambochila.

Drosophila montgomeryi, north Kaluaa Gulch.

Drosophila montgomeryi North Kaluaa 4596
Drosophila montgomeryi. North Kaluaa Gulch.

Drosophila montgomeryi North Kaluaa 4598
Drosophila montgomeryi. North Kaluaa Gulch.

Drosophila montgomeryi North Kaluaa 4611
Drosophila montgomeryi. North Kaluaa Gulch.

Drosophila montgomeryi on a rotting Urera kaalae branch, north Kaluaa Gulch.

Drosophila montgomeryi on a rotting Urera kaalae branch, north Kaluaa Gulch....
Drosophila montgomeryi on a rotting Urera kaalae branch, north Kaluaa Gulch. Note the eggs laid in the bark on the middle left (the white threads are respiratory filaments).

Drosophila montgomeryi on a rotting Urera kaalae branch, north Kaluaa Gulch....
Drosophila montgomeryi on a rotting Urera kaalae branch, north Kaluaa Gulch. These two males are getting ready to fight.

Drosophila montgomeryi on a rotting Urera kaalae branch, north Kaluaa Gulch....
Drosophila montgomeryi on a rotting Urera kaalae branch, north Kaluaa Gulch. Note the eggs laid in the bark near where it's feeding (the white threads are respiratory filaments).

Eupithecia sp Kaluaa 6126
Eupithecia sp., feeding on Drosophila ambochila. North Kaluaa Gulch,...
Eupithecia sp., feeding on Drosophila ambochila. North Kaluaa Gulch, Honouliuli. We've all seen pictures of one eating a Drosophila in the lab, but I've always wondered if they really did it in the wild. Now we know...

Eupithecia sp Kaluaa 6128
Eupithecia sp., feeding on Drosophila ambochila. North Kaluaa Gulch,...
Eupithecia sp., feeding on Drosophila ambochila. North Kaluaa Gulch, Honouliuli. We've all seen pictures of one eating a Drosophila in the lab, but I've always wondered if they really did it in the wild. Now we know...

Eupithecia sp Kaluaa 6131
Eupithecia sp., feeding on Drosophila ambochila. North Kaluaa Gulch,...
Eupithecia sp., feeding on Drosophila ambochila. North Kaluaa Gulch, Honouliuli. We've all seen pictures of one eating a Drosophila in the lab, but I've always wondered if they really did it in the wild. Now we know...

Aloha nr plectranthi Hapapa 3886
Aloha nr. plectranthi. Puu Hapapa.

Drosophila ambochila Hapapa 4383
February 23, 2010. Puu Hapapa shelf. Drosophila ambochila.

Drosophila ambochila Hapapa 4387
February 23, 2010. Puu Hapapa shelf. Drosophila ambochila.

Drosophila ambochila Hapapa 4388
February 23, 2010. Puu Hapapa shelf. Drosophila ambochila.

Drosophila crucigera Hapapa 4405
February 23, 2010. Puu Hapapa shelf. Drosophila crucigera.

Drosophila crucigera Hapapa 4406
February 23, 2010. Puu Hapapa shelf. Drosophila crucigera.

Drosophila crucigera Hapapa 4580
February 25, 2010. Puu Hapapa shelf. Drosophila crucigera.

Drosophila inedita Hapapa 4396
February 23, 2010. Puu Hapapa shelf. Drosophila inedita.

Drosophila inedita Hapapa 4603
February 25, 2010. Puu Hapapa shelf. Drosophila inedita.

Drosophila inedita Hapapa 4605
February 25, 2010. Puu Hapapa shelf. Drosophila inedita.

Drosophila montgomeryi Hapapa 4473
February 24, 2010. Puu Hapapa shelf. Drosophila montgomeryi. Distinguished...
February 24, 2010. Puu Hapapa shelf. Drosophila montgomeryi. Distinguished from D. ambochila by the two brown stripes on the thorax, the distinct dark line on the upper aneipsternum, the elongate, dark cerci, and the multiple rows of long hairs on the front tibia.

Drosophila montgomeryi Hapapa 4475
February 24, 2010. Puu Hapapa shelf. Drosophila montgomeryi. Distinguished...
February 24, 2010. Puu Hapapa shelf. Drosophila montgomeryi. Distinguished from D. ambochila by the two brown stripes on the thorax, the distinct dark line on the upper aneipsternum, the elongate, dark cerci, and the multiple rows of long hairs on the front tibia.

Drosophila montgomeryi Hapapa 4482
February 24, 2010. Puu Hapapa shelf. Drosophila montgomeryi. Distinguished...
February 24, 2010. Puu Hapapa shelf. Drosophila montgomeryi. Distinguished from D. ambochila by the two brown stripes on the thorax, the distinct dark line on the upper aneipsternum, the elongate, dark cerci, and the multiple rows of long hairs on the front tibia.

February 24, 2010. Puu Hapapa shelf. Drosophila montgomeryi. Distinguished from D. ambochila by the two brown stripes on the thorax, the distinct dark line on the upper aneipsternum, the elongate, dark cerci, and the multiple rows of long hairs on the front tibia. All of these are visible in this picture.

February 25, 2010. Puu Hapapa shelf. Drosophila montgomeryi. Note the two...
February 25, 2010. Puu Hapapa shelf. Drosophila montgomeryi. Note the two brown stripes on the thorax, compared to the completely pale thorax of D. ambochila on the left.

February 25, 2010. Puu Hapapa shelf. Drosophila montgomeryi. Note the two...
February 25, 2010. Puu Hapapa shelf. Drosophila montgomeryi. Note the two brown stripes on the thorax.

Drosophila punalua Hapapa 4397
February 23, 2010. Puu Hapapa shelf. Drosophila punalua.

Drosophila punalua Hapapa 4398
February 23, 2010. Puu Hapapa shelf. Drosophila punalua.

Drosophila punalua Hapapa 4401
February 23, 2010. Puu Hapapa shelf. Drosophila punalua.

Drosophila spp Hapapa 4590
February 25, 2010. Puu Hapapa shelf. Drosophila spp.: D. crucigera on the...
February 25, 2010. Puu Hapapa shelf. Drosophila spp.: D. crucigera on the upper right, D. inedita lower left, D. montgomeryi upper center, and three D. ambochila across the middle.

caterpillar Hapapa 4449
February 23, 2010. Puu Hapapa shelf.

caterpillar Hapapa 4450
February 23, 2010. Puu Hapapa shelf.

caterpillar Hapapa 4452
February 23, 2010. Puu Hapapa shelf.

cockroach Hapapa 4438
February 23, 2010. Puu Hapapa shelf. Native(?) cockroach.

cockroach Hapapa 4444
February 23, 2010. Puu Hapapa shelf. Native(?) cockroach.

grasshopper eating snail Hapapa 4440
February 23, 2010. Puu Hapapa shelf. Grasshopper eating a small snail.

Nesophrosyne Hapapa 4434
February 23, 2010. Puu Hapapa shelf. Nesophrosyne sp.

Orthotylus sp Hapapa 4432
February 23, 2010. Puu Hapapa shelf. Orthotylus sp.

Prognathogryllus sp Hapapa 9848
Prognathogryllus sp., Hawaiian tree cricket. Puu Hapapa, found by Jamie Tanino.

Prognathogryllus sp Hapapa 9852
Prognathogryllus sp., Hawaiian tree cricket. Puu Hapapa, found by Jamie Tanino.

Prognathogryllus sp Hapapa 9855
Prognathogryllus sp., Hawaiian tree cricket. Puu Hapapa, found by Jamie Tanino.

Taeniaptera angulata Hapapa 4494
February 24, 2010. Puu Hapapa shelf. Taeniaptera angulata (Micropezidae).

Taeniaptera angulata Hapapa 4495
February 24, 2010. Puu Hapapa shelf. Taeniaptera angulata (Micropezidae).

Vanessa tameamea Hapapa 4497
February 24, 2010. Puu Hapapa shelf. Vanessa tameamea, Kamehameha butterfly.

Vanessa tameamea Hapapa 4500
February 24, 2010. Puu Hapapa shelf. Vanessa tameamea, Kamehameha butterfly.

Vanessa tameamea Hapapa 4501
February 24, 2010. Puu Hapapa shelf. Vanessa tameamea, Kamehameha butterfly.

February 24, 2010. Puu Hapapa shelf. Vanessa tameamea, Kamehameha butterfly...
February 24, 2010. Puu Hapapa shelf. Vanessa tameamea, Kamehameha butterfly caterpillar on Urera.

Zaprionus indianus Hapapa 9984
Zaprionus indianus, Puu Hapapa. It's very distinctive with white stripes...
Zaprionus indianus, Puu Hapapa. It's very distinctive with white stripes extending all along the body. This widespread tramp species was only discovered in Hawaii a few months ago, but it already occurs throughout the Waianae range, sometimes as only a few individuals and sometimes in large numbers like this.

Drosophila ambochila. Puu Hapapa, Honouliuli.

Drosophila gradata Hapapa 9568
Drosophila gradata. Puu Hapapa, Honouliuli.

Drosophila larifuga Hapapa 9589
Drosophila larifuga on rotting Urera kaalae bark. Puu Hapapa, Honouliuli.

Drosophila larifuga Hapapa 9593
Drosophila larifuga on rotting Urera kaalae bark. Puu Hapapa, Honouliuli.

Drosophila montgomeryi, Puu Hapapa. They are often able to detect the sound...
Drosophila montgomeryi, Puu Hapapa. They are often able to detect the sound of a shutter button and take off as the picture is taken.

Drosophila montgomeryi, Puu Hapapa. A courtship pileup!

Enicospilus sp Hapapa 9604
Enicospilus sp. Puu Hapapa, Honouliuli. These native nocturnal wasps were...
Enicospilus sp. Puu Hapapa, Honouliuli. These native nocturnal wasps were aggregating inside in large numbers during the day, something I hadn't seen before.

Neomachilis insularis Hapapa 3899
Neomachilis insularis, the native jumping bristletail. Puu Hapapa.

Neomachilis insularis Hapapa 3901
Neomachilis insularis, the native jumping bristletail. Puu Hapapa.

Neomachilis insularis Hapapa 3902
Neomachilis insularis, the native jumping bristletail. Puu Hapapa.

oribatid mite Hapapa 3909
Oribatid mite on Pisonia branch. Puu Hapapa.

oribatid mite Hapapa 3910
Oribatid mite on Pisonia branch. Puu Hapapa.

Drosophila montgomeryi courting, Puu Hapapa.

Drosophila hemipeza Hapapa 5232
Drosophila hemipeza, Puu Hapapa. This is the first sighting of this species...
Drosophila hemipeza, Puu Hapapa. This is the first sighting of this species outside of Palikea since 1975.

Drosophila hemipeza Hapapa 5233
Drosophila hemipeza, Puu Hapapa. This is the first sighting of this species...
Drosophila hemipeza, Puu Hapapa. This is the first sighting of this species outside of Palikea since 1975.

Drosophila hemipeza Hapapa 5234
Drosophila hemipeza, Puu Hapapa. This is the first sighting of this species...
Drosophila hemipeza, Puu Hapapa. This is the first sighting of this species outside of Palikea since 1975.

Drosophila montgomeryi. Puu Hapapa.

Drosophila montgomeryi courting on a bait sponge, Puu Hapapa.

Drosophila montgomeryi males competing on a bait sponge, Puu Hapapa.

Drosophila montgomeryi Moho Gulch 5386
Drosophila montgomeryi, Moho Gulch, Honouliuli.

Drosophila montgomeryi Moho Gulch 5391
Drosophila montgomeryi, Moho Gulch, Honouliuli.

Drosophila montgomeryi Moho Gulch 5393
Drosophila montgomeryi, Moho Gulch, Honouliuli.

Drosophila sp Pleomele Oio 3972
Drosophila sp., Oio Gulch, Kahuku. This is a modified mouthparts species...
Drosophila sp., Oio Gulch, Kahuku. This is a modified mouthparts species that breeds in the leaf axils of Pleomele, feeding on the detritus that accumulates there. Even when it's quite dry the center of the axil retains water. They then pupate higher up on the leaves.

Drosophila sp Pleomele Oio 3973
Drosophila sp., Oio Gulch, Kahuku. This is a modified mouthparts species...
Drosophila sp., Oio Gulch, Kahuku. This is a modified mouthparts species that breeds in the leaf axils of Pleomele, feeding on the detritus that accumulates there. Even when it's quite dry the center of the axil retains water. They then pupate higher up on the leaves.

Drosophila sp Pleomele Oio 3974
Drosophila sp., Oio Gulch, Kahuku. This is a modified mouthparts species...
Drosophila sp., Oio Gulch, Kahuku. This is a modified mouthparts species that breeds in the leaf axils of Pleomele, feeding on the detritus that accumulates there. Even when it's quite dry the center of the axil retains water. They then pupate higher up on the leaves.

Drosophila obatai Palikea gulch 9654
Drosophila obatai. Palikea Gulch below Puu Pane, Kaala NAR.

Drosophila obatai Palikea gulch 9657
Drosophila obatai. Palikea Gulch below Puu Pane, Kaala NAR.

Drosophila obatai Palikea gulch 9659
Drosophila obatai. Palikea Gulch below Puu Pane, Kaala NAR.

Drosophila substenoptera Kalena 4584
Drosophila substenoptera, in South Haleauau below Puu Kalena.

Drosophila obatai Pulee 5702
Drosophila obatai, Pulee Gulch (Guava Gulch), Schofield Barracks.

Drosophila obatai Pulee 4221
Drosophila obatai. Pulee Gulch, above Schofield Barracks.

Drosophila obatai Pulee 4222
Drosophila obatai. Pulee Gulch, above Schofield Barracks.

Drosophila obatai. Pulee Gulch, above Schofield Barracks.

Drosophila oahuensis North Haleauau 5120
Drosophila oahuensis, North Haleauau Gulch.

Drosophila oahuensis North Haleauau 5121
Drosophila oahuensis, North Haleauau Gulch.

Lasioglossum impavidum Manuwai 2461
Lasioglossum impavidum, an introduced sweat bee now widespread around Hawaii....
Lasioglossum impavidum, an introduced sweat bee now widespread around Hawaii. Manuwai ridge.

Drosophila pilimana Manuwai 3860
Drosophila pilimana. Manuwai Gulch, Kaala NAR.

Drosophila pilimana Manuwai 3853
Drosophila pilimana. Manuwai Gulch, Kaala NAR.

Drosophila obatai Manuwai 1036
Drosophila obatai, Manuwai Gulch.

Drosophila paucicilia Manuwai 1132
Drosophila paucicilia, Manuwai Gulch.

Drosophila pilimana Manuwai 1123
Drosophila pilimana, Manuwai Gulch.

Drosophila pilimana, Manuwai Gulch.

Drosophila reynoldsiae Manuwai 1029
Drosophila reynoldsiae, Manuwai Gulch.

Drosophila reynoldsiae Manuwai 1030
Drosophila reynoldsiae, Manuwai Gulch.

Drosophila reynoldsiae Manuwai 1031
Drosophila reynoldsiae, Manuwai Gulch.

Hyposmocoma sp Manuwai 5185
A very camoflaged case of a Hyposmocoma caterpillar on bark of a lama tree....
A very camoflaged case of a Hyposmocoma caterpillar on bark of a lama tree. Manuwai Gulch.

Vanessa tameamea Manuwai 5184
Kamehameha butterflies, Vanessa tameamea, on Drosophila bait. Just as I...
Kamehameha butterflies, Vanessa tameamea, on Drosophila bait. Just as I snapped a picture of one feeding on the fermented banana, another flew right in front.

Drosophila flexipes Manuwai 5151
Drosophila flexipes, Manuwai Gulch.

Drosophila flexipes Manuwai 5152
Drosophila flexipes, Manuwai Gulch.

Drosophila obatai Manuwai 3841
Drosophila obatai? Manuwai Gulch, Kaala NAR.

Drosophila reynoldsiae? Manuwai Gulch, Kaala NAR.

Drosophila sp Manuwai 5153
Drosophila sp., Manuwai Gulch. This species superficially resembles D....
Drosophila sp., Manuwai Gulch. This species superficially resembles D. turbata but has the scutellum dark.

Drosophila spp Manuwai 5143
Drosophila flexipes (left) and D. gradata (right), Manuwai Gulch.

Drosophila obatai Manuwai 5149
Drosophila obatai, Manuwai Gulch.

Drosophila flexipes Manuwai 1047
Drosophila flexipes, Manuwai Gulch.

Drosophila obatai, Manuwai Gulch.

Drosophila paucicilia Manuwai 1077
Drosophila paucicilia, Manuwai Gulch.

Drosophila paucicilia Manuwai 5161
Drosophila paucicilia, Manuwai Gulch.

Drosophila sp Manuwai 1085
Drosophila sp., Manuwai Gulch. This species superficially resembles D....
Drosophila sp., Manuwai Gulch. This species superficially resembles D. turbata but has the scutellum dark.

Drosophila sp Manuwai 1086
Drosophila sp., Manuwai Gulch. This species superficially resembles D....
Drosophila sp., Manuwai Gulch. This species superficially resembles D. turbata but has the scutellum dark.

Drosophila spp Manuwai 1082
Drosophila obatai (left) and D. flexipes (right), Manuwai Gulch.

Drosophila spp Manuwai 5165
Drosophila flexipes (top and bottom) and D. gradata (middle), Manuwai Gulch.

Drosophila crucigera (above), D. paucicilia (below center), and D. flexipes...
Drosophila crucigera (above), D. paucicilia (below center), and D. flexipes (right), Manuwai Gulch.

Drosophila paucicilia and flexipes Manuwai 3862
Drosophila paucicilia (left) and D. flexipes (right), both very rare breeders...
Drosophila paucicilia (left) and D. flexipes (right), both very rare breeders on sap fluxes of Sapindus oahuensis (lonomea). Manuwai Gulch, Kaala NAR.

Hylaeus anthracinus Kahuku 3679
Hylaeus anthracinus on Heliotropium foertherianum (=Tournefortia argentea)....
Hylaeus anthracinus on Heliotropium foertherianum (=Tournefortia argentea). Kahuku Point, Oahu.

Hylaeus anthracinus Kahuku 3680
Hylaeus anthracinus on Heliotropium foertherianum (=Tournefortia argentea)....
Hylaeus anthracinus on Heliotropium foertherianum (=Tournefortia argentea). Kahuku Point, Oahu.

Hylaeus anthracinus Kahuku 3681
Hylaeus anthracinus on Heliotropium foertherianum (=Tournefortia argentea)....
Hylaeus anthracinus on Heliotropium foertherianum (=Tournefortia argentea). Kahuku Point, Oahu.

Hylaeus longiceps Kahuku 3684
Hylaeus longiceps on Heliotropium foertherianum (=Tournefortia argentea)....
Hylaeus longiceps on Heliotropium foertherianum (=Tournefortia argentea). Kahuku Point, Oahu.

Hylaeus longiceps Kahuku 3685
Hylaeus longiceps on Heliotropium foertherianum (=Tournefortia argentea)....
Hylaeus longiceps on Heliotropium foertherianum (=Tournefortia argentea). Kahuku Point, Oahu.

Hylaeus longiceps Kahuku 3687
Hylaeus longiceps on Heliotropium foertherianum (=Tournefortia argentea)....
Hylaeus longiceps on Heliotropium foertherianum (=Tournefortia argentea). Kahuku Point, Oahu.

Hylaeus longiceps Kahuku 3687a

Drosophila deltaneuron Kaala 5135
Drosophila deltaneuron, just off the summit of Kaala. This unusual modified...
Drosophila deltaneuron, just off the summit of Kaala. This unusual modified mouthparts species has wing markings very similar to many species in the picture wing group.

Drosophila nigribasis Kaala 5134
Drosophila nigribasis, just off the summit of Kaala. This male would...
Drosophila nigribasis, just off the summit of Kaala. This male would normally have the darker pattern of the right wing, but for some reason the color only developed on one side.

Drosophila nigribasis Kaala 5138
Drosophila nigribasis, just off the summit of Kaala. This is a female,...
Drosophila nigribasis, just off the summit of Kaala. This is a female, distinguished by the less extensive wing coloration.

Drosophila nigribasis Kaala 5139
Drosophila nigribasis, just off the summit of Kaala. This is a female,...
Drosophila nigribasis, just off the summit of Kaala. This is a female, distinguished by the less extensive wing coloration.

Drosophila oahuensis Kaala 4566
Drosophila oahuensis. Kaala.

Drosophila nigribasis, female. Kaala.

Drosophila neogrimshawi Kaala 9860
Drosophila neogrimshawi, Kaala. This rare species has not been seen since...
Drosophila neogrimshawi, Kaala. This rare species has not been seen since 1972 and apparently not in the Waianae range since 1916. At Kaala, it lives on the very steep slopes and gulches below the summit bog where there are stands of Clermontia, its host plant.

Drosophila neogrimshawi Kaala 9864
Drosophila neogrimshawi, Kaala. This rare species has not been seen since...
Drosophila neogrimshawi, Kaala. This rare species has not been seen since 1972 and apparently not in the Waianae range since 1916. At Kaala, it lives on the very steep slopes and gulches below the summit bog where there are stands of Clermontia, its host plant.

Drosophila neogrimshawi Kaala 9866
Drosophila neogrimshawi, Kaala. This rare species has not been seen since...
Drosophila neogrimshawi, Kaala. This rare species has not been seen since 1972 and apparently not in the Waianae range since 1916. At Kaala, it lives on the very steep slopes and gulches below the summit bog where there are stands of Clermontia, its host plant.

Drosophila obatai. Manuwai Gulch.

Telostylinus lineolatus Manuwai 4201
Telostylinus lineolatus, an introduced stilt-legged fly. Manuwai Gulch.

Sierola Kaala 1201
May 26, 2007. Kaala near the twin towers, Oahu. Sierola (Bethylidae) on the...
May 26, 2007. Kaala near the twin towers, Oahu. Sierola (Bethylidae) on the underside of a leaf. Dan Polhemus calls this area "Hunter's forest"; it has a reasonably tall canopy with moss-covered trunks, unlike the boggy, more open forest around the boardwalk.

Sierola Kaala 1202
May 26, 2007. Kaala near the twin towers, Oahu. Sierola (Bethylidae) on the...
May 26, 2007. Kaala near the twin towers, Oahu. Sierola (Bethylidae) on the underside of a leaf. Dan Polhemus calls this area "Hunter's forest"; it has a reasonably tall canopy with moss-covered trunks, unlike the boggy, more open forest around the boardwalk.

Waianae-Kaala Trail, just below Kaala summit plateau. Drosophila nigribasis...
Waianae-Kaala Trail, just below Kaala summit plateau. Drosophila nigribasis (male).

Drosophila nigribasis, male. Kaala.

Waianae-Kaala Trail, just below Kaala summit plateau. Drosophila nigribasis...
Waianae-Kaala Trail, just below Kaala summit plateau. Drosophila nigribasis (female).

Drosophila oahuensis Kaala 7977
Waianae-Kaala Trail, just below Kaala summit plateau. Drosophila oahuensis...
Waianae-Kaala Trail, just below Kaala summit plateau. Drosophila oahuensis (female).

Drosophila oahuensis Kaala 7980
Waianae-Kaala Trail, just below Kaala summit plateau. Drosophila oahuensis...
Waianae-Kaala Trail, just below Kaala summit plateau. Drosophila oahuensis (male).

Waianae-Kaala Trail, just below Kaala summit plateau. Drosophila oahuensis...
Waianae-Kaala Trail, just below Kaala summit plateau. Drosophila oahuensis (male).

Waianae-Kaala Trail, just below Kaala summit plateau. Drosophila oahuensis (male).

Waianae-Kaala Trail, just below Kaala summit plateau. Drosophila oahuensis (female).

Drosophila substenoptera Kaala 7996
Waianae-Kaala Trail, just below Kaala summit plateau. Drosophila substenoptera.

Drosophila substenoptera Kaala 7998
Waianae-Kaala Trail, just below Kaala summit plateau. Drosophila substenoptera.

Drosophila substenoptera Kaala 7999
Waianae-Kaala Trail, just below Kaala summit plateau. Drosophila substenoptera.

Drosophila pilimana Kaala 7968
Waianae-Kaala Trail, just below Kaala summit plateau. Drosophila pilimana.

Drosophila pilimana Kaala 7975
Waianae-Kaala Trail, just below Kaala summit plateau. Drosophila pilimana.

Drosophila pilimana Kaala 7976
Waianae-Kaala Trail, just below Kaala summit plateau. Drosophila pilimana.

Drosophila punalua Kaala 7973
Waianae-Kaala Trail, just below Kaala summit plateau. Drosophila punalua.

Drosophila punalua Kaala 7974
Waianae-Kaala Trail, just below Kaala summit plateau. Drosophila punalua.

Drosophila spp Kaala 7961
Waianae-Kaala Trail, just below Kaala summit plateau. Drosophila punalua...
Waianae-Kaala Trail, just below Kaala summit plateau. Drosophila punalua (above) and D. pilimana (below).

Drosophila spp Kaala 7965
Waianae-Kaala Trail, just below Kaala summit plateau. Drosophila punalua...
Waianae-Kaala Trail, just below Kaala summit plateau. Drosophila punalua (right) and D. pilimana (left).

Nesorestias sp Kaala 1197
May 26, 2007. Kaala boardwalk, Oahu. Nesorestias sp., or possibly...
May 26, 2007. Kaala boardwalk, Oahu. Nesorestias sp., or possibly Nothorestias. Neal pointed this out to me, saying it was one of the flightless dolichopodids (Emperoptera) that hang around on Melicope leaf litter and noting how surprising it was that it was letting me take the picture without fleeing. I commented on how short and squat it was, and that I would have thought it was a delphacid if he hadn't pointed it out. Turns out there was a reason for that.

Drosophila kinoole Waianae 0926
Drosophila kinoole, Waianae Valley. This is only the second of this species...
Drosophila kinoole, Waianae Valley. This is only the second of this species ever seen; it was previously known from a single teneral, poorly-preserved specimen reared in 1971, so its appearance wasn't even completely known.

Drosophila kinoole Waianae 0928
Drosophila kinoole, Waianae Valley. This is only the second of this species...
Drosophila kinoole, Waianae Valley. This is only the second of this species ever seen; it was previously known from a single teneral, poorly-preserved specimen reared in 1971, so its appearance wasn't even completely known.

Drosophila kinoole Waianae 0929
Drosophila kinoole, Waianae Valley. This is only the second of this species...
Drosophila kinoole, Waianae Valley. This is only the second of this species ever seen; it was previously known from a single teneral, poorly-preserved specimen reared in 1971, so its appearance wasn't even completely known.

Drosophila montgomeryi Waianae 5110
Drosophila montgomeryi on bait sponge, Waianae Valley.

Drosophila hexachaetae Waianae 1143
Drosophila hexachaetae, Waianae Valley.

Drosophila kinoole, Waianae Valley.

Drosophila montgomeryi Waianae 1137
Drosophila montgomeryi, Waianae Valley. Two males preparing to fight for a...
Drosophila montgomeryi, Waianae Valley. Two males preparing to fight for a lek spot.

Drosophila montgomeryi Waianae 1138
Drosophila montgomeryi, Waianae Valley. Two males preparing to fight for a...
Drosophila montgomeryi, Waianae Valley. Two males preparing to fight for a lek spot.

Drosophila montgomeryi, Waianae Valley. Two males attempting to court the...
Drosophila montgomeryi, Waianae Valley. Two males attempting to court the same female.

Drosophila montgomeryi ovipositing in a beetle borehole in Urera glabra,...
Drosophila montgomeryi ovipositing in a beetle borehole in Urera glabra, Waianae Valley.

Drosophila montgomeryi and a native Tetragnatha spider, Waianae Valley.

Drosophila pilimana Waianae 5533
Drosophila pilimana, Waianae Valley.

Laupala sp Waianae 5574
Laupala sp., Waianae Valley.

Drosophila montgomeryi, Waianae Valley.

Hyposmocoma habitat Waianae 5771
August 8, 2010. Waianae Valley, first gulch north of Puu Kawiwi. Kumaipo...
August 8, 2010. Waianae Valley, first gulch north of Puu Kawiwi. Kumaipo Stream, where we collected the aquatic Hyposmocoma.

Hyposmocoma hunting Waianae 5769
August 8, 2010. Waianae Valley, first gulch north of Puu Kawiwi. Akito...
August 8, 2010. Waianae Valley, first gulch north of Puu Kawiwi. Akito Kawahara collecting semi-aquatic Hyposmocoma caterpillars from rocks.

Hyposmocoma hunting Waianae 5770
August 8, 2010. Waianae Valley, first gulch north of Puu Kawiwi. Akito...
August 8, 2010. Waianae Valley, first gulch north of Puu Kawiwi. Akito Kawahara collecting semi-aquatic Hyposmocoma caterpillars from rocks.

Hyposmocoma Waianae burrito 5764
August 8, 2010. Waianae Valley, first gulch north of Puu Kawiwi....
August 8, 2010. Waianae Valley, first gulch north of Puu Kawiwi. Hyposmocoma sp., one of the semi-aquatic species. The stream appeared to be long dry, but the caterpillars were abundant, presumably still grazing on the rocks. This "burrito case" species was found higher on the rocks and in drier spots than the sympatric "cone case" species.

Hyposmocoma Waianae burrito 5765
August 8, 2010. Waianae Valley, first gulch north of Puu Kawiwi....
August 8, 2010. Waianae Valley, first gulch north of Puu Kawiwi. Hyposmocoma sp., one of the semi-aquatic species. The stream appeared to be long dry, but the caterpillars were abundant, presumably still grazing on the rocks. This "burrito case" species was found higher on the rocks and in drier spots than the sympatric "cone case" species.

Hyposmocoma Waianae burrito 5767
August 8, 2010. Waianae Valley, first gulch north of Puu Kawiwi....
August 8, 2010. Waianae Valley, first gulch north of Puu Kawiwi. Hyposmocoma sp., one of the semi-aquatic species. The stream appeared to be long dry, but the caterpillars were abundant, presumably still grazing on the rocks. This "burrito case" species was found higher on the rocks and in drier spots than the sympatric "cone case" species.

Hyposmocoma Waianae cone 5759
August 8, 2010. Waianae Valley, first gulch north of Puu Kawiwi. Hyposmocoma sp., one of the semi-aquatic species. The stream appeared to be long dry, but the caterpillars were abundant, presumably still grazing on the rocks. This "cone case" species was found only in damp spots and was much less common than the sympatric "burrito case" species.

Lepidopsocus Waianae 5760
August 8, 2010. Waianae Valley, first gulch north of Puu Kawiwi. Lepidopsocus sp. These were common in the streambed, running around on the rocks.

Lepidopsocus Waianae 5761
August 8, 2010. Waianae Valley, first gulch north of Puu Kawiwi. Lepidopsocus sp. These were common in the streambed, running around on the rocks.

Hylaeus anomalus Kumaipo 3366
Hylaeus anomalus, Kamaileunu ridge. This species has not been seen since...
Hylaeus anomalus, Kamaileunu ridge. This species has not been seen since 1930 except for two specimens in 1977, and never before in the Waianae range.

Hylaeus anomalus Kumaipo 3368
Hylaeus anomalus, Kamaileunu ridge. This species has not been seen since...
Hylaeus anomalus, Kamaileunu ridge. This species has not been seen since 1930 except for two specimens in 1977, and never before in the Waianae range.

Drosophila pilimana Makaha 0002
Drosophila pilimana, Makaha Valley.

Drosophila pilimana Makaha 4726
Drosophila pilimana, Makaha Valley.

Drosophila pilimana Makaha 4728
Drosophila pilimana, Makaha Valley.

Hylaeus connectens Makaha 2164
September 25, 2009. Makaha Valley, Chamaesyce herbstii exclosure on south...
September 25, 2009. Makaha Valley, Chamaesyce herbstii exclosure on south wall. Hylaeus connectens on Claoxylon sandwicense.

Hylaeus connectens Makaha 2166
September 25, 2009. Makaha Valley, Chamaesyce herbstii exclosure on south...
September 25, 2009. Makaha Valley, Chamaesyce herbstii exclosure on south wall. Hylaeus connectens on Claoxylon sandwicense.

Hylaeus connectens Makaha 2167
September 25, 2009. Makaha Valley, Chamaesyce herbstii exclosure on south...
September 25, 2009. Makaha Valley, Chamaesyce herbstii exclosure on south wall. Hylaeus connectens on Claoxylon sandwicense.

September 25, 2009. Makaha Valley, Chamaesyce herbstii exclosure on south wall. Hylaeus connectens on Chaemaesyce herbstii.

Scaptomyza palata Makaha 5803
August 9, 2010. Makaha Valley. Scaptomyza (Grimshawomyia) palata.

Scaptomyza palata Makaha 5806
August 9, 2010. Makaha Valley. Scaptomyza (Grimshawomyia) palata.

Scaptomyza palata Makaha 5807
August 9, 2010. Makaha Valley. Scaptomyza (Grimshawomyia) palata.

Drosophila spaniothrix Makaleha 1882
Drosophila spaniothrix, Central Makaleha Gulch. This is the first record of...
Drosophila spaniothrix, Central Makaleha Gulch. This is the first record of this species since 1971.

Drosophila spaniothrix Makaleha 1883
Drosophila spaniothrix, Central Makaleha Gulch. This is the first record of...
Drosophila spaniothrix, Central Makaleha Gulch. This is the first record of this species since 1971.

Drosophila spaniothrix Makaleha 1889
Drosophila spaniothrix, Central Makaleha Gulch. This is the first record of...
Drosophila spaniothrix, Central Makaleha Gulch. This is the first record of this species since 1971.

Drosophila obatai Makaleha 5654
Drosophila obatai, East Makaleha Gulch, west fork.

Drosophila obatai Makaleha 5656
Drosophila obatai, East Makaleha Gulch, west fork.

Drosophila sobrina Makaleha 5661
Drosophila sobrina. East Makaleha Gulch, west fork.

Drosophila sobrina Makaleha 5663
Drosophila sobrina. East Makaleha Gulch, west fork.

Eupithecia sp West Makaleha 4152
Eupithecia sp., carnivorous caterpillar. West Makaleha Gulch, Kaala.

Eupithecia sp West Makaleha 4154
Eupithecia sp., carnivorous caterpillar. West Makaleha Gulch, Kaala.

Drosophila turbata Ohikilolo 9534
Drosophila turbata. Ohikilolo.

Drosophila turbata Ohikilolo 9535
Drosophila turbata. Ohikilolo.

Drosophila turbata Ohikilolo 9536
Drosophila turbata. Ohikilolo.

Zaprionus sp Ohikilolo 9549
Zaprionus sp. Ohikilolo.

Zaprionus sp Ohikilolo 9541
Zaprionus sp. Ohikilolo.

Nysius sp Ohikilolo 9530
Nysius sp., native seed bugs feeding on ohia inflorescences. Ohikilolo.

Nysius sp Ohikilolo 9532
Nysius sp., native seed bugs feeding on ohia inflorescences. Ohikilolo.

Drosophila ant predation Ohikilolo 4147
Big-headed ants (Pheidole megacephala) tearing apart a native picture wing...
Big-headed ants (Pheidole megacephala) tearing apart a native picture wing fly, Drosophila punalua. Ohikilolo. Despite the ants' presence, there was a moderate number of flies here including one rare species.

Drosophila crucigera Ohikilolo 9520
Drosophila crucigera. Ohikilolo.

Vanessa tameamea predation Kapuna 1691
Evident predation by birds on Kamehameha butterfly (Vanessa tameamea),...
Evident predation by birds on Kamehameha butterfly (Vanessa tameamea), Mokuleia Trail, Pahole.

Nesiomiris timberlakei Kapuna 1681
Nesiomiris timberlakei on Reynoldsia (Polyscia) sandwicensis. Mokuleia...
Nesiomiris timberlakei on Reynoldsia (Polyscia) sandwicensis. Mokuleia Trail, Pahole.

Nesiomiris timberlakei Kapuna 1683
Nesiomiris timberlakei on Reynoldsia (Polyscia) sandwicensis. Mokuleia...
Nesiomiris timberlakei on Reynoldsia (Polyscia) sandwicensis. Mokuleia Trail, Pahole.

Nesiomiris timberlakei Kapuna 1685
Nesiomiris timberlakei on Reynoldsia (Polyscia) sandwicensis. Mokuleia...
Nesiomiris timberlakei on Reynoldsia (Polyscia) sandwicensis. Mokuleia Trail, Pahole.

Goniozus sp Pahole 4535
Goniozus sp., Pahole Gulch branch 3. This introduced species is relatively...
Goniozus sp., Pahole Gulch branch 3. This introduced species is relatively widespread but is unidentified.

Goniozus sp Pahole 4558
Goniozus sp., Pahole Gulch branch 3. This introduced species is relatively...
Goniozus sp., Pahole Gulch branch 3. This introduced species is relatively widespread but is unidentified. While resting on leaves bethylids will often bend their abdomen around and twist onto their side; it's not clear why.

Goniozus sp Pahole 4560
Goniozus sp., Pahole Gulch branch 3. This introduced species is relatively...
Goniozus sp., Pahole Gulch branch 3. This introduced species is relatively widespread but is unidentified. While resting on leaves bethylids will often bend their abdomen around and twist onto their side; it's not clear why.

Sierola sp Pahole 4543
Sierola sp., Pahole Gulch branch 3.

Sierola sp Pahole 4544
Sierola sp., Pahole Gulch branch 3.

Sierola sp Pahole 4550
Sierola sp., Pahole Gulch branch 3. While resting on leaves bethylids will...
Sierola sp., Pahole Gulch branch 3. While resting on leaves bethylids will often bend their abdomen around and twist onto their side; it's not clear why.

Sierola sp., Pahole Gulch branch 3. While resting on leaves bethylids will...
Sierola sp., Pahole Gulch branch 3. While resting on leaves bethylids will often bend their abdomen around and twist onto their side; it's not clear why.

Drosophila montgomeryi larva Hapapa 5070
Drosophila montgomeryi eggs and larva in rotting bark of Urera kaalae. At...
Drosophila montgomeryi eggs and larva in rotting bark of Urera kaalae. At least 28 eggs are visible in this picture; they were almost certainly laid all at once by one female.

Dictyophorodelphax mirabilis Pahole 4918
Dictyophorodelphax mirabilis, the akoko planthopper. The gut extends into...
Dictyophorodelphax mirabilis, the akoko planthopper. The gut extends into the long horn and may be used to detoxify the milky sap of its host plant.

Dictyophorodelphax mirabilis Pahole 4920
Dictyophorodelphax mirabilis, the akoko planthopper. The gut extends into...
Dictyophorodelphax mirabilis, the akoko planthopper. The gut extends into the long horn and may be used to detoxify the milky sap of its host plant.

Dictyophorodelphax mirabilis Pahole 4925
Dictyophorodelphax mirabilis, the akoko planthopper. The gut extends into...
Dictyophorodelphax mirabilis, the akoko planthopper. The gut extends into the long horn and may be used to detoxify the milky sap of its host plant.

black widow Peacock Flat 4846
March 25, 2010. Peacock Flat campground. Black widow?

Heteropoda venatoria Peacock Flat 4904
March 27, 2010. Peacock Flat campground toilet. Heteropoda venatoria, cane...
March 27, 2010. Peacock Flat campground toilet. Heteropoda venatoria, cane spider.

Heteropoda venatoria Peacock Flat 4906
March 27, 2010. Peacock Flat campground toilet. Heteropoda venatoria, cane...
March 27, 2010. Peacock Flat campground toilet. Heteropoda venatoria, cane spider.

Heteropoda venatoria Peacock Flat 4909
March 27, 2010. Peacock Flat campground toilet. Heteropoda venatoria, cane...
March 27, 2010. Peacock Flat campground toilet. Heteropoda venatoria, cane spider.

Scolopendra subspinipes Peacock Flat 4840
March 25, 2010. Peacock Flat campground. Scolopendra subspinipes.

Scolopendra subspinipes Peacock Flat 4841
March 25, 2010. Peacock Flat campground. Scolopendra subspinipes.

Scolopendra subspinipes Peacock Flat 4842
March 25, 2010. Peacock Flat campground. Scolopendra subspinipes.

Hylaeus strenuus Keaau 8969
Hylaeus strenuus, a recently-introduced bee from India, on Erythrina...
Hylaeus strenuus, a recently-introduced bee from India, on Erythrina sandwicensis, wiliwili. Keaau Valley, Makua-Keaau FR. This is a typically bird-pollinated flower, and no other bees here have been seen visiting it.

Hylaeus strenuus Keaau 8972
Hylaeus strenuus, a recently introduced bee from India. Keaau Valley,...
Hylaeus strenuus, a recently introduced bee from India. Keaau Valley, Makua-Keaau FR. It is blowing out a drop of wiliwili (Erythrina sandwicensis) nectar to concentrate it before returning to its nest. Erythrina are bird-pollinated and have relatively dilute nectar, meaning it's inefficient for them to carry it without concentrating. No other bees here seem to visit it.

Hylaeus strenuus Keaau 8972b
Hylaeus strenuus, a recently introduced bee from India. Keaau Valley,...
Hylaeus strenuus, a recently introduced bee from India. Keaau Valley, Makua-Keaau FR. It is blowing out a drop of wiliwili (Erythrina sandwicensis) nectar to concentrate it before returning to its nest. Erythrina are bird-pollinated and have relatively dilute nectar, meaning it's inefficient for them to carry it without concentrating. No other bees here seem to visit it.

Hylaeus strenuus Keaau 8973
Hylaeus strenuus, a recently introduced bee from India. Keaau Valley,...
Hylaeus strenuus, a recently introduced bee from India. Keaau Valley, Makua-Keaau FR. It is blowing out a drop of wiliwili (Erythrina sandwicensis) nectar to concentrate it before returning to its nest. Erythrina are bird-pollinated and have relatively dilute nectar, meaning it's inefficient for them to carry it without concentrating. No other bees here seem to visit it.

Hylaeus strenuus Keaau 8976b

Quadrastichus erythrinae Keaau 8962
Quadrastichus erythrinae, erythrina gall wasp, on flowers of wiliwili...
Quadrastichus erythrinae, erythrina gall wasp, on flowers of wiliwili (Erythrina sandwicensis). Keaau Valley, Makua-Keaau FR. This area formerly had a dense grove of trees, but many were killed by the wasp. Even though it is largely controlled now, many of the surviving trees are mostly dead, and the wasp continues to cause damage to flowers and seed pods.

Quadrastichus erythrinae Keaau 8963
Quadrastichus erythrinae, erythrina gall wasp, on flowers of wiliwili...
Quadrastichus erythrinae, erythrina gall wasp, on flowers of wiliwili (Erythrina sandwicensis). Keaau Valley, Makua-Keaau FR. This area formerly had a dense grove of trees, but many were killed by the wasp. Even though it is largely controlled now, many of the surviving trees are mostly dead, and the wasp continues to cause damage to flowers and seed pods.

Quadrastichus erythrinae Keaau 8964
Quadrastichus erythrinae, erythrina gall wasp, on flowers of wiliwili...
Quadrastichus erythrinae, erythrina gall wasp, on flowers of wiliwili (Erythrina sandwicensis). Keaau Valley, Makua-Keaau FR. This area formerly had a dense grove of trees, but many were killed by the wasp. Even though it is largely controlled now, many of the surviving trees are mostly dead, and the wasp continues to cause damage to flowers and seed pods.

Jadera haematoloma Kealia 2863
Jadera haematoloma, the alien red-shouldered soapberry bug, feeding on seeds...
Jadera haematoloma, the alien red-shouldered soapberry bug, feeding on seeds of Sapindus oahuensis. Kealia Trail.